... 09◦00’ W 08◦52’ WAltitude (m) 20 0–3 00 20 0–4 00 0–1 00 0–1 00Mean annual rainfall (mm) 70 0–8 00 70 0–1 000 50 0–6 00 50 0–8 00Mean temperature (◦C) 12. 5–1 5 1 5–1 6 1 5–1 6 1 5–1 7Soil type Eutric litosoils ... y−1).Within the site, the trees also showed differences in growth,especially in diameter, with coefficients of variation of the mean around 20% (Tab. II). The extent and the pattern of the within-tree ... account the presence of heartwood in the trees and the factors of its variation.Acknowledgements: We thank StoraEnso for allowing the sam-pling, Jorge Gominho for advice in heartwood determinations,...