đại học cần thơ đại học cần thơ đại học cần thơ đại học cần thơ - -- - khoa nông nghiệp khoa nông nghiệp khoa nông nghiệp khoa nông nghiệp giáo trình giảng dạy trực tuyến QJ7S&ĂQ7K7HO)D[ :HEVLWHKWWSZZZFWXHGXYQNQQHPDLOGPLQK#FWXHGXYQYWDQK#FWXHGXYQ Giáo trình Các nguyên lý về bệnh hại cây trồng Chơng 9 Các biện pháp không dùng hóa chất Trong phòng trị bệnh hại cây trồng Biên soạn bởi PGs. Phạm Văn Kim Bộ môn Bảo Vệ Thực Vật Khoa Nông Nghiệp Đại Học Cần Thơ &DÉF1JX\½Q/\ÉY½Ð%½ÕQK+DÕL&»\7U¾ÐQJ&KÀ¿QJ &+ml1* &$±&%,¥½13+$±3.+¦1*'8®1*+2±$&+£¼7 7521*3+2®1*75%¥½1++$½,&£<75¦¸1* iSKRQJWUEQKKDLF\WUQJFKXQJWDFQDSGXQJWWFDNLQWKFYEQK F\DKRFSKLKSYLNLQWKFYQQJKRFYORDLF\LWQJORDLF\FQ EDRYNFDFDFNLQWKFYFDFODQKYFFROLQTXDQQKNLQKWWKWUQJPL WUQJ YY PLFRWKDWFNWTXDPRQJPXQ7X\QKLQGWPKLX FKXQJWD[SFDFELQSKDSFRWKDSGXQJYDRFDFQKRPNKDFQKDXWX\WKHRWQK FKWFXDWQJELQSKDS&DFQKRPELQSKDSJPFRELQSKDSFDQKWDFELQ SKDSVLQKKRFELQSKDSO\KRFYDFKRFELQSKDSKRDKRFELQSKDSNLPGFK WKF YW YD JQ \ ELQ SKDS TXDQ O\ GFK KDL PW FDFK WQJ KS ,30 LQWHJUDWHGSHVWVPDQDJHPHQWFXQJFQKFQKQQKLX ,%,j13+$v3&$1+7$v& 0XFFK 7DRLXNLQWKXQOLFKRVSKDWWULQFXDF\WUQJF\WUQJFR XVFFKQJLODLYLKXKWFDFEQK 7DRLXNLQWKXQOLFKRFDFYLVLQKYWLNKDQJSKDWWULQWWF FKVSKDWWULQFXDPPEQK 7DRLXNLQNKQJWKXQKSFKRPPEQKYFDFPWSKDWWULQWFK OX\YDODQWUX\Q &DFELQSKDSFDQKWDFFRKLXTXDWURQJSKRQJQJDEQKF\ D/DPW iWODQLOXWQFXDUWQKLXORDLPPEQK'RRWFKQKODQJXQG WUWFKOX\YDO\ODQFXDEQK.KLFD\EDWFKXQJWDODPWKD\LO\WQKFX WUXFPQKLWFXDW\WRODPWKD\LLXNLQVQJYDSKDWWULQFXD PPEQK.KLFD\WFKXQJWDYXLPWVPPEQK[XQJVXGLWODPFKR FKXQJFKWKRFNKRNKQWURQJKRDWQJJ\KDLFKRF\ &DÉF1JX\½Q/\ÉY½Ð%½ÕQK+DÕL&»\7U¾ÐQJ&KÀ¿QJ &D\EDWFRQWDRLXNLQWKRDQJNKFKRWJLXSVFKX\QKRDWURQJ WF[D\UDWWKQWSKRQJWKFKGQJOLXPWFDFKLXKRDKQJLXSFKR F\WUQJSKDWWULQWWWEPFEQKKQ &D\EDWFRQJLXSVFKX\QKRDFDF[DFEDWKFYWWWKQFXQJFS QKLXQQJOQJFKRKYLVLQKYWVQJWURQJWQKWODFDF[DNKXQYDFDFYL VLQKYWNKDFFRNKDQQJLNKDQJYLPPEQK 'RRVDXNKL[RQJPWYXFKXQJWDFQSKDLFD\WSKLDLQXFRWKL JLDQ&RQQXYPXDYXNKWNKDRTXDWKWQKWFXQJSKDLFD\EDWPWOQPL QP 9LFFD\DLSKLWWURQJPWWKLJLDQQKWQKWURQJQPFRDQKKQJ NKDTXDQWURQJLYLPWVEQKF\7KGXQKWURQJWUQJKSFXDEQK YDQJODYDEQKFKD\ODOXDWDLi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ÉF1JX\½Q/\ÉY½Ð%½ÕQK+DÕL&»\7U¾ÐQJ&KÀ¿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i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ÉF1JX\½Q/\ÉY½Ð%½ÕQK+DÕL&»\7U¾ÐQJ&KÀ¿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ÉF1JX\½Q/\ÉY½Ð%½ÕQK+DÕL&»\7U¾ÐQJ&KÀ¿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ÉF1JX\½Q/\ÉY½Ð%½ÕQK+DÕL&»\7U¾ÐQJ&KÀ¿QJ JLXSF\FRXVFFKQJFKXYLEQKYDJLXSF\VPKLSKXFVDXWGFK EQKQQJ7X\QKLQQXFKXQJWDODPGXQJSKQKRDKRFERQYLOLXOQJOQ TXDKRFERQTXDQKLXDPPWFDFKPWFQLVHDQKXTXDQJFYL PRQJPXQODF\VHPFEQKQQJODPJLDPQQJVXWNKQJNLQKW &KXQJWDELWOD 3KQDPTXDFDRODPJLDPGD\FXDOSFXWLQEDRFKHELXEODODP FKRODGQKLPEQKKQ 3KQOQJLXSWQJGD\FXDOSFXWLQJLXSSKQQDRWURQJVFKQJ FKXYLEQKFXDF\WUQJltWSKHQF\WKLXOQWUPWURQJQQPFXFKQJ EQKYDQQJVXWFXQJUWNHP%RQOQFKRF\WURQJWUQJKSQ\JLXSF\ SKDWWULQWWWQJWQKFKQJEQKPWFDFKURUWYDWQJQQJVXWYDSKP FKWDQJN 3KQNDOLJLXSWQJWQKFKQJEQKFXDF\WUQJ.KLF\PFEQK QQJVDXNKLDSGXQJFDFELQSKDSWFKFFFKQJEQKSKXQWKXFYY QQERQWKPPWOQJNDOLQKWQKVHJLXSF\FKQJEQKWWKQYDF\PDX KLSKXFKQ7X\QKLQPWVWUQJKSNKDFYLFFRKD\NKQJFRERQSKQ NDOLFXQJNKQJDQKKQJJOQEQKFD7'EQKYDQJODOXDltWUQJKSNKDF QDWKDQKKQJFXDSKQ.OQEQKSKFWDSKQQKLX7'EQKFKD\ODOXDGR 3\ULFXODULDRU\]DH &DFORDLYLOQJSKQOQQXFR\XWURQJF\VHFRWDFGXQJJLXS F\FKQJEQKWWKQ&XQJFSYLOQJFKRF\EQJFDFKSKXQGXQJGFKYL OQJOQOD ,,%,j13+$v36,1++2& iLQKQJKDYDPXFFKFXDELQSKDSVLQKKRF %LQ SKDS VLQK KRF WURQJ SKRQJ WU EQK F\ ODLXNKLQPLWUQJF\ WUQJYDVLQKYWLNKDQJPWFDFKWKFKKSWDRQQPWWKFQEQJVLQK KRFFQWKLWJLXSJLDPPWVFXDPPEQK[XQJGLQJQJJ\KDL1KR EQKFXDF\WUQJFK[XWKLQPFQKHNKQJJ\DQKKQJTXDQWURQJY PWNLQKW &DÉF1JX\½Q/\ÉY½Ð%½ÕQK+DÕL&»\7U¾ÐQJ&KÀ¿QJ %LQ SKDSVLQKKRFNKQJFRPXFFKWLXGLWWRDQEPPEQKYDFXQJ NKQJFRNKDQQJQ\ 7WFDFDFELQSKDSLSKRYLEQKF\FRDQKKQJOQKDLQKRPVLQKYW FXQJPWOXFF\WUQJYDPPEQKYLVLQKYWLNKDQJYDPPEQKF\ WUQJYDYLVLQKYWLNKDQJYYXFOLWYDRELQSKDSVLQKKRF &QEQJVLQKKRFWURQJWKLQQKLQ D7KJLLVLQKKRFYDFQEQJVLQKKRFWURQJWKLQQKLQ 7KJLLVLQKKRFODPWPDQJOLUQJOQWURQJRFDFFGQFXDVLQKYW OXQOXQWDFQJOQQKDXWDRQQPWWKFQEQJi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hQiDWDRUDFDFGFKEQKFKD\EDODOXDWUP WURQJWURQJQKLXQP 'FKEQKPFVQJNKRDLW\WDL,UHODQGGRVGXQJQJORDWJLQJ NKRDLW\QKLPEQK &DÉF1JX\½Q/\ÉY½Ð%½ÕQK+DÕL&»\7U¾ÐQJ&KÀ¿QJ lt9LW1DPYLFVGXQJWUQGLQUQJFDFJLQJOXDQKLPU\QXWURQJ FDFQPYDDDQGFKU\QXWUPWURQJWDLi%6&/ 6GXQJFDFORDLWKXFWUVXTXDFFDQJODPFKRU\QXJLDWQJ9DRFXQJOD PWWURQJFDFQJX\QQKQJ\QQGFKEQKOXQ[RQODOXDWDLFDFWQK&Q 7K6RF7UQJ9QK/RQJ7UD9LQKYDiQJ7KDSYDRYXQJ[XQ 7UQ X QDQK YLF SKXQ WKXF WU VX ED EDLODPFKRGFK VX [DQK 6SRGRSWHUDH[LJXDQJD\FDQJWUPWURQJWKPYDVX[DQKQJD\FDQJNKDQJWKXF KQ ls3KDSYLFQKSJLQJQKRWFKX0\YFXQJQJWKLHPWKHRPP EQK3ODVPRSDUDYLWLFRODDWLXGLWWRDQEYQQKRFXD3KDSWURQJQKLXQP OLQ 1JD\QD\FKXQJWDELWUWFKFFKQUQJFDFGFKEQKWUPWURQJ[D\UDOD GRFRQQJLDEQJQKLXFDFKSKDYFQEQJVLQKKRF;D[DVXEQK[D\UD YLPFQKHJ\KDLNKQJDQJNQKELWDFQJFXDFRQQJLOQK VLQKWKDLFKDOQFKDJ\QQVPWFQEQJVLQKKRFTXDQWURQJQKQJD\ QD\ltWKLN\Q\PPEQKGXFRPWFXQJNKQJJ\KDLTXDWUPWURQJY FRQEFDFVLQKYWNKDFFFK %LQSKDSVLQKKRFWURQJEQKF\ D&DF\XWVGXQJWURQJELQSKDSVLQKKRF 7URQJELQSKDSVLQKKRFFKXQJWDFQWPKLXWDFQJOQQKDXFXDFDF QKRPVLQKYWYDWDFQJFXDPLWUQJOQWQJQKRPWKXF\VSKDWWULQ WFKFFFXDPWYDLQKRPFXQJQKFFKFDFWDFQJWLXFFFXDPPEQK&DF QKRPVLQKYWPDFKXQJWDFRWKVGXQJWURQJELQSKDSVLQKKRFODN\FKXPP EQKYDYLVLQKYWLNKDQJ1JRDLUDPLWUQJODPW\XWKWVFTXDQWURQJ WURQJELQSKDSVLQKKRF α αα α.\FKX 7URQJWQKWUDQJWQKLQNKQJFRFRQQJLWDFQJYDRN\FKXOXQOXQ WURQJWQKWUDQJFEDRYELFQEQJVLQKKRFWURQJPLWUQJVQJFXD FKXQJ1JX\QQKQFRWKOD &DÉF1JX\½Q/\ÉY½Ð%½ÕQK+DÕL&»\7U¾ÐQJ&KÀ¿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mvQJGXQJJLDWKX\WQ\KLQQD\FR QKLXQKDNKRDKRFWSWUXQJQJKLQFXFDFKNFKWKFKWQKNKDQJEQKFXDFDF JLQJ QKLPEQKWUFNKLF\EPPEQKWQFQJ6NFKWKFKWQKNKDQJ EQKQ\FWKFKLQEQJFDFKSKXQKRDFKWNKQJSKDLODWKXFEDRYWKF YWKRFSKXQPWORDLYLVLQKYWNKQJJ\KDLFKRF\\+RDFKWKRFYLVLQK YWNFKNKDQJVHWDFQJOQWKXWKQJRDLPWODYDNFKWKFKFDFWKXWKQ\ &DFWKXWKENFKWKFKVHWUX\QFDFWQKLXYDRWURQJWEDRNFKQJFDFJLHQ WLWUDFKWNKDQJEQKKRDWQJWUFNKLEPPEQKWQFQJiQNKLPP EQKWQFQJWURQJWEDRDFRVQFDFFKWNKDQJYLPPEQK\1KRF\ WKRDWEQKKRFPFEQKQKHKQ%LQSKDSQ\FJRLODNFKWKFKWQKNKDQJ EQKFXDF\WUQJLQGXFHGUHVLVWDQFH &R KDL ORDL NFK WKFK WQK NKDQJ EQK NFK WKFK WQK NKDQJ WDLFKORFDO LQGXFHGUHVLVWDQFHYDNFKWKFKWQKNKDQJOXGQV\VWHPLFDFTXLUHGUHVLVWDQFH %LQ SKDS NFKWKFKWQKNKDQJOXGQFDFKXQJKQYDFWSWUXQJ QJKLQFXQKLXKQ . :HEVLWHKWWSZZZFWXHGXYQNQQHPDLOGPLQK#FWXHGXYQYWDQK#FWXHGXYQ Giáo trình Các nguyên lý về bệnh hại cây trồng Chơng 9 Các biện pháp không dùng hóa chất Trong phòng trị bệnh hại cây trồng Biên soạn bởi PGs.