Nghiên cứu đặc điểm hình thái hóa sinh và sự đa dạng di truyền của một số giống lúa cạn địa phương

!" # $ % &' ! & ( ) / 12 ) * ,$ & )+ , ,- ! &4 $ , )9 1; < ,$ & ," / ! = $ ,E ? F J BB : >? @ L H = AB D: )K B : = C 4B D & H # )K L= M N / ! : % ! A) ? = I L= I AO P # %? $ I % &' A >? @ $ ! %? & = " >? @ G $ B ! Q2 I # DK # SSSSSSSSSSSSSSSSSSSSSSS DK $ SSSSSSSSSSSSSSSSSSSSSSSS T2 SSSSSSSSSSSSSSSSSSSSSSSSS R B U W U & " V SSSSSSSSSSSSSSSSSSSSS Q Q SSSSSSSSSSSSSSSSSSSSS X ,$ SSSSSSSSSSSSSSSSSSSSS Y $ %&'((((((((((((((((((((((((( ) *+ ,, , 34 B 1'((((((((((((( ,, SSSSSSSSSSSSSSSSSSSSSS ,, - '/ Z & ? L[ \ $ 78 ? G 55 ? G ,> ? , C DE = SSSSSSSSSSSSSSSS = SSSSSSSSSSSS R " S SSS ^ 57 ((((((((((( ,= = SSSSSSSSSSSSSS _ R ,6 > - > : = ; < = ? @ 78 B ` SSSSSSSSSSSSSSS "7 F/ G 7H & ] A 34 ((((((((( ? @ S R' M0 V = L[ / R5 SSSSSSSSSSSSSSSSSS Q R R R_ U 5Z J" # $ ? R SSSSSSS ? ? h !" # $ & F > DX ' > , %9 R gfMQSSSSSSSSSSSSSSSS \ Rd `SSSSSSSSSSS R5 55R^ M = 5Y CSSSSSSS R M g I gfMQ SSSSSSSSSSSSSSSS 55RR M $ 5Y SSSSSSSS 5Y ? G ? ? ? 5X SSSSSSSSSSSS 5Y c SSSSSSSSSSSSSSSS )+ M )8 = SSSSSSSSSS 6W ` SSSSSSSSSSSSSSSSSS 5Y ? 5 ^ M )8 55R @ (((((((((((((((( ? ?? ? )8 555 ? V< 57 A V [ 34 >S > = SSSSSSSSSSSS RW )+ = SSSSSSSSSSS Ra ? RX ? ` = ^ )K SSSSSS > = S ^d R _ M N > %9 ? : R5 ` = U \ @ A ]: ^ V [ > = 34 S [ L_ ` 5((((((((((((((((((((( i = & R55 L < ! $ ' > > Da7 b \ U 78 4 )+ K A V ? [ 34 ` @ U 7c((((((((((((((((((((((( RR M F RR5 T ( # fQB ,j / !k U a % gfMQSSSSSSS > = 1( C> = )8 = ^5 ? CW ^X F gfMQSSSSSSSSSSSSSSSSSSSSSSSSSS DX =W J" J" 'N M0 %d e ((((((((((((((((( % SSSSSSSSSSSSSSSSSSSSSSS SSSSSSSSSSSSSSSSSSSSSSSS 1' / D _X _Y Y(((((((((((((((((( Sf " ] SSSSSSSSSSSSSSSSSSSSS Wd " f SSSSSSSSSSSSSSSSSSSSS W5 _ g fHf f, fQB f fmDM , ?` ` % o? - p rggr r ` J, J , ` JQ J Dqf D ` q , M g M ` gfMQ g & ` ` ` ` f r l ! g` & ? & ` D` '( ; g, ` ` A Q 1` A ? M )8 ? ? " #$ ` ) If ` ` I qQ f q t qQ f l2 ' ! ` W 34 % s '- s ! % : M ? l2 & " #$ ` ? fq , - / QBf l2 & ( ? ` A`# ` ` g`?` ` ? l2 ) , f ? n `1 M m ` l* + / % '( % A ) f, '( ` g` AQAIMfPq ? & g ` g` ` -4 AQA M - ` `Q - ` D` ( q gmDM ` '( qQ f AAg QBf l & E gB I ` f ? n `1 n fA h f 1` QBf gM X c AQA G/ # P ) \ 6, ? )8 \ 66 ? G R \ > , J" # $ ? ^ \ > L[ a _ \ > a \ = >= >B b E & ! ` & )K = O RW )+ ddd = Ra = = RY ? ` = a ^ < )+ = O = l s ? _ > ,, b > & = )9 & ! N )8 ? = >) > ,f = > ^_ = ^W = ? p gfMQ &9 _ \ R = = \ ? )+ R _ a \ ? i Y ? > 5W = >S \ = a )+ >C b \ d ` ? = = X > > > >> b \ Z \ > W & Z ? Z )8 h ,O a ; ; a ^a = = fQB )+ = fQB Z C ,$ & ? $ gfMQ ! $ a > / a ^Y _^ = > = G/ # P i ! , , A8 Z 6, b R 66T ^ >,M N _ >6 W >> b &9 a $ = c & # # ? ` > = = = ! = > a h ,O $ ? p gfMQ >= b 5X > ^R = ^^ a > = $ $ ? p gfMQ a > $ $ ? p gfMQ a > $ $ ? p gfMQ a > $ ? p gfMQ a > # / Z Ta >S b Z T R >B b = &9 $ Z T ^ > W A8 Z > &/ = X 5a / Z T^ = &9 d ! >C b = &9 Y $ > Z TR = &9 X $# U _d _d _ _5 _R _W $ %&' %j Mk %d D> )8 -? )8 ( G )K 12 , ( # O > / O )8 )+ Z r % ` " )8 G ,V = )! ' ) - " ' - ! p = = V B )+ ! $ )+ = &9 - ^5 ' : O 5dd5 [ \ !k * E )K D Q - * ' YXY ] B' YYY ' c? ? G \ > ' YY_ B - ! p ' O F ' )+ R W )9 & * - ! p " _l = - # ?& " ,E ! > ! )+ " )9 )+ ? " ` vaw \ A &9 -? [ ,j "? & * 5a i ! V? x * Z B = aI du ? 1` s B K ? 12 B Z / &9 ( ? )K ] ] )+ )9 5dd5 : = RR W - &9 ^ _W - ! p - > 5ddR c ! $ IRu ? : &)8 - ! p / = )9 )9 G - > > O] ( = 5_ c Ydu z LE H / Z B' > O ! V? > )K F V ,c &9 G y " ( = > A$ / K o D> > & E = )9 B = - )9 vaw { &9 > )9 )K = U &{ " a _u ? F ,V l D> ! c ? N ! - )+ = c / = > " " F Z > HV s &{ / [ F , + ) &4 F ! ! c! ' Q b $ # h ) ! $ ' \ ` Y K -? )8 $ )9 M c ? c c? ? G / ! HV L = > ( Q F , O Z < HE B = | > / > S = ` Z O U 8 1} ? { +? &9 " )9 " )K # h 1{ vYw F = ch ~ > b \ ! - )+ G > = G = ? )8 & ! $ ' 5dd5 v5^w• B C e - ? * : U V< & A - )+ = E 55 L > = ! $ ' = & M Q RR M F R^ M E E ? & ? G ? \ > b C J 7H ` c - b$ €" T% 5dd^ v aws : " F )K / x ? @ A l7 Z c & - ! p ' F = > & ," : c O? 1( U / ' > c : E & ! $ ' = = ? & ,$ C > c E )+ ( / )K c %? ( ? > & &% > = < > = = l = h c > = % ? = fQB * ` > = gfMQ &9 Z > = " %? d ; # \ / x TR T^ Ta T R T ^ J" # $ E \ >) ? _ R E *^ " E * ? - d? - ? = # $? s -? F gfMQ _ \ O ? = H5 c !F / R Ys = gfMQ i u? Xb ,$ O Z x W‹W ? = = 5_ d ^^ ^ Rd d dd d 5_ d [bEb ^ R W = F ,$ & ? M u? | Z ! ^_ _u TZ T R c " x ‹^ ? ddu = " Z TR & T ^ c 5_u Q)9 !" Z TR T^ Ta T R & T ^ $ ? p R d L c gfMQ )9 F cR? F )+ )+ c )9 d R J, ƒ !F ? a ,$ * d R J, " = fQB )+ bP AD5 AD - > = fQB )+ / ? gfMQ $ ? p *R " ^? = l,$ = - / ,\ > J" # $ D i Z = fQB )+ M = = ! ? $ ? = " W? Z ; = fQB ,$ ^ Y d W ^ ZZ TR T^ Ta T R T ^ $_ = fQB )+ Z M H$ - U ! TZ % ,$ )8 )9 d aW J, = fQB ^Y Z TR _ J, c / H5 > &9 JF - ^ - )9 D & H ,$ š &9 > D & Z TR )+ ; = fQB )+ \ = &c - R J, Z TR a ? _ J, - = $ > = &4 F c > _ J, R J, d aWJ, d RJ, >>b $ $ ? p gfMQ =O s > = &9 Z TR e C J" # $ \ a O $ ? p gfMQ > = &9 Z T^ )+ R^ _ J, J, d X J, d R J, >C b $ $ ? p gfMQ =O a e _d > = &9 Z T^ TZ T^ \ F *W " Y? = > &9 fQB R Y? bP AD5 > D ? ? ! / = fQB AD5 H5 - ^ J, J, ! = fQB D ! - H5 AD X? = H R ^s L / bP AD5 l = - / - H5 D AD - )8 ? - c (! _ J, ! ( ) &% H D bP ? = fQB O - &/ [ / = &9 Z Ta )+ U !\ B O $ ? p gfMQ > R_ d J, d J, d W J, d R J, >= b $ $ ? p gfMQ =O e _ a < = fQB ! = fQB O &4 F d _R J, ! AD = &4 F - - = d J, d X J, d ^_ = fQB = &4 F d RJ, Œ J" # $ \ )9 a = fQB ? D ? > s W? ; _ J, H5 & D = &4 F d W5 J, c ^ = &4 F d _R J, - - ? = fQB - D - - = fQB )+ )9 * d R J, " = fQB ƒ !F J, - $ c c !F Z T^ c R H - ? > = &9 Z Ta A ? ? = fQB = &9 !F = fQB )+ - )9 ! \ ? bP AD5 Sb $ $ ? p gfMQ =O a e _5 > = &9 Z T R J" # $ - RW ? ? = fQB = fQB B )+ - - ? s - c ( ! )8 = fQB ? &/ = &9 Z T ^ )+ Ra Y J, ^ J, d a J, d _ J, >Bb $ $ ? p gfMQ =O M F !" # $ - ? = > = &9 Z T ^ e $ ? p *R " ^? c a / )9 F gfMQ &9 Z T ^ = fQB )+ * d _ J, " _R ,$ Y J, a ; * !" # $ > Ra - ! cR? \ - F ? ? F = &9 !F Z E? J" # $ : )+ # ; U a $ ? p gfMQ X5 ? = fQB | - = fQB )+ ,$ F ? = fQB )+ ,$ N ? \ > ,f ! &/ [ \ O ,$ = &9 _ Z ; R d $ 5W ? = - Rd ? = - P D c AD5 c 5^ ? = TZ Ta c N - _Y ? = TZ TR & T R c ,$ F d > - 5R ? ` : = - B ) &% c (! 12 U a = N ? _ P \ / = fQB )+ / - B ) &% !" # $ ( a ! ? c - Y J, > ,$ ; / T ^ : = fQB )+ > bP & AD5 ! ! ? Z ^ J, = fQB / O _ d _ J, d a J, & = &4 F gfMQ &9 H )9 )9 F = fQB Z - ; ! a $ ? p > gfMQ &9 = TR T^ Ta T R T ^ N ^ X X ^ 5W H5 ^ X a R ^ 5W H R W Y ^ ^ 5W D R X d _ ^ Rd bP R W Y R 5R AD5 R W Y R R 5^ AD R Y a ^ ^ 5a YZ bX b_ YZ Yd XcY D _^ >>6 [ b T / U - )8 > &9 \ D G F & d lf?? `1 H - # = s &/ r / [ [? / c )8 )8 Z Z / -? > Z H5 / H a > D = bP AD5 AD dddd H5 d XaXX dddd H d a_aW d X X5 dddd D d WYWY d WYWY d X X5 dddd bP d a5aR d a5aR d X^X_ d a5aR dddd AD5 d WYWY d a_aW d XaXX d a_aW d YWYa dddd AD d X^X_ d YdY d X^X_ d aXaY d WYWY d a5aR T E! )+ , \ U )+ )+ = fQB gfMQ )+ •dŠ! > )8 D - / ? $ / > ,, b P F F/ G E? ? ? )+ = O - gfMQ A & JI 7H? U 78 > )9 x Š b # 34 B A€A? &` Z > [ ! )8 YYXs l ` # b U $ ? p ,j „Af • B ( ; ! F K # " %? 1( ? T J 7H "? & E > = c PU )8 H$ R * d WYWY " d YWYa ,j ! c Z - c = Z c ( / ! / • &/ E &9 / )8 [? )8 dddd Z D & D / * D & AD5 [? j O! H5 & D $ bP & AD c / )8 - c [? J" # $ ? O Z F # RX - -? - ld WYWYs )8 Z = )+ \ bP & AD5 ld YWYas F RX b / Z - a R & / U > > )+ c Z ^ c &9 F ! 5Wu l dd I d a^ Š d 5Ws > W A8 Z B c c D F &/ - & # Z R / U > )+ c H5 AD &9 = < ! R _u l dd t d XW_ Š d R_s B c c D F Z & cR ^ 5R 5u l dd t d aWX Š d 5R5s B & AD5 c ! -? )+ H c5 - lR dus _W < bP AD5 &9 ? ! c bP b c i bP & AD5 c ' )+ &/ "? E ! $ ' ? = ! F = )+ ? ! c c )8 - ? , O -? $ < = X R^ & = Z _a = / Yau P &9 # R Y ^WWs - l^ _ G & ^ 5Y G s E 5Ru B ) &% fQB & ! $ ' l U D ! &9 (? > = c c G O )+ "? DX 'N M0 %d e , Da7 " , , L: ? = a ^ E > = &9 > ? 2K # & R E 1E * a RY ddd = / > c - )+ ,6 b = )+ P c b )+ )+ = ? = )K - )+ ? - J bG " * -? )+ WRu " ^ a_u - " H5 l WRus a_ P > AD l d s b = c H lY us M -? ' E d c c )+ c ? ` u ` > c )8 > * H5 l a_s " 5X R AD l^ a_us a > }• ? 2J > - )K u a ? = * Y u " & = )+ ,> b = "? W )+ ` > ? ` - 1( U = E * Y u I AD5 l us -? = :,\ = > - F L: )+ ( c > ` ? ` ` E ,C = = ! = "l ` )+ ! < ,4 = O = - l < = a ? , S Hj !k % gfMQ : B' = &9 )8 = )+ = = s b c (? $ ! > = ` ' a = & N * a _a u I d _u > = a c ca = ? [ Z )+ & b = = ( ,\ O i N )+ bP & AD5 c ! $ ' )8 X R^ & R Y ^WW• i ^ _ G & ^ 5Y G s ,$ )+ 12 _X X5 ? = fQB * ` / ,\ F F )+ c ddu ? = ,B Z D > R lD s &9 - O Z T^ Ta T R lTZ T R s = ? H5 AD s & c ! Z \ , Z 5Wu b K Œ c c F ^ c - l H bP AD5 bP & AD5 c ! $ ' )8 Z / - lYaus lbP AD5s c \ 12 = 6%K > c! $ ' = &% G = c U > &/ = c? ? G 3 _Y c ` # > c! $ ' " ! $ ' 4 0 1' D X G H '( ) M = 12 b BE R B D ^ M / D Y M1 D i - T E l YYXs & -g - E BeH L= # & A l Yaas C bU )K B C G H l5ddRs •T O > ‚ ( 3$ h #L b BE 4J f )8 # + $ , V ( B M '% ? ` # h B C b)8 i b €" l5dd s •B h b BE W IWX V BeH P 4T E & F ; " 3$ , j ! l E D % &' B G Q Q)8 D) B b / ]Œ L y T l YYYs •A0 12 ? )8 ? ? gfMQ 17 > ? &2 ‚ mM h '$ / b BE ^_5I ^_a a D Q : D D HF D e b)8 B C T B ‚ mM )' / L 4M D U )+ ? h E l s X_IYR ]' & A l YYas •A0 12 E gfMQ O > Q IRR & ‚ ( _I X _ L| Q)8 l5dd s '% b - )K L= A) ? = W 1' \ #^ %E = ~ A E^ -k 3$ b " E V + b L| > ] - )+ X D e LV L 4M D 4T E D G H l YYYs •A0 12 !k % gfMQ \ F fBQ E 17 3 * ‡ > a ‚ mM V +I h '$ / b BE R^ I R^a Y B C b b T% & A l5ddRs •L 1= = O / > ? F HV ] B ‚ n 3i " ' R ' h Eb " XWIXY Wd , > l ) d B L C [? d I dX h b= M = l5dd^s •B = ' G 4b 1= Rd_I R y b 1= - ‚ " T B C E 17 > + h Yl s / =? ` D l YYYs •A0 12 / O ( &% ‚ mM !k % M gIgfMQ VF +I h '$ / E J/ b)8 l5ddRs , j V 3$ o i M 0 - ( 3^ p Y``Y ( % E D % &' = ~B B D B R rggr l YYWs m+ M ? ` e % % g f ER ) o M K K ' ? )K L= & - T ^ M = B l5dd^s , ( ! %E $ q '% ( 3$ #( # " M 0 - ( f r P; Q( 3$ F ' P\ D % &' = ~A )K L= A) ? = B B _ b t W T% l5dd s s , M 0 - ( V ' 3p u P; * + j l '% 3$ h U q h -h v I L/ B]Jb -? ,E & - b T% B 4T D e LV L 4M D G H l5ddds •œ 12 !k % ? F )+ ? = fBQ )+ ,$ ; lgfMQs & & ,E ` 17 % E ," ‚ ( I h lRs _I5R a b ? + X L ' ( T% l5dd^s •B ` = E , l ) ' R ' " E B 4M D G H = > O # " 3$ , #^ &/ ? ` & ? G > = ? )8 ‚ n 3i h E V -& _ aI_5d D = R W T E l YYWs •e = ,j ? )8 ? ? _IX = " ! $ = ‚ Y L 4M D G B C ]' ] " c F ! ! ,j !k % gfMQ‚ _a I_a^ 5d B C L %? ? F HV ] G ]' = &{ h ] b " T B bZ l YYXs •L = ? )8 & %? E O < / > # " 3$ , #^ l R s I_ > B ‚ i B B + 55 D) B o‚ F IR 5R B , B = H D 4T E B C b$ b D Q l5ddRs •B 1= %? > &9 , ,= & ! p w ' ' ' n 3i " , l ) ' R ' h ( C ‚E 5I W l YYYs •J %' R ' h l YYas •M = + h 3$ , C b$ - B + ' ? b BE J J -& 5^ B C b$ €" l5dd5s I ( 3$ j ) , l E D % &' = ~ A B B 1' X #^ ` 1` A€ n x > r & ˆ? @X 3$ Y PM ` ( > h EI # > Z + 3$ R R1 / ) % E BeH B g , ( ! % E '% R7 M 0 - ( V )K L= A) M = / l YaWs • 1 &` n nf 5a % l YYWs wt -k + 5_ H ` D A l YaRs •g ? 1` ` ` • 2I' -RYx 5d_ I 5da 5W Z `‘ Ÿ Db n ` ` n n `` ? `& fž ` & `• y AD `• W5 `n ” ` I Q M ^R_I^^ l YY s •T ` , P' RR ? _aR I _WY ` 5X ` , A b T B l5dd5s •q ` , ` `` n ,` ` 2P '- bO 5_dI5_a 5Y Q M ` @A y Rd rggr R J 2- R5 D ` E YYa D` T DC M ? 5Ya I RdR - ` ` n ` D` H 1` &`1 n ED H M P b l YYas •‘ ?` ='- P 'E R_ ? 5_ I R^ ˆ J l Yads R_ ’` , l5dd s •gfMQ & 1` I ` ?? ` M 55 I 5_ & fQB & , • n `• 55a M WXdIWXX RR T D T f ! ! Ž D m ! gfMQ !` ` ` &` lA A s A A I A?` n ? ` • R^ g` m D H g D A b n ` ` ` ? fQB AAg !` • ' ` ` n , ` ? ,` `• F A l YY^s •Q` ` `# ` ` XE ?? RY I ^X ˆ Ÿ € l5ddRs •f ? ` ff I ` ` ‘ Ž ?` ` -G E dX l s R I 5d ˆ T ` T l YYds •m ` ? ` ? ` • B f g` X a5 R I a5 X ` RW ’ ˆ P J J ,` f q D`& ! J ˆ g n ž ! ˆ f l YYds •QBf ? ? ? n `1 , , ? ` ` ` !` • B f g` X l55s W_R I W_R_ WR n `1 ` &` gfMQ M g” ` A] ` `n ... "? ! l aI5 ,' / dI W JQ E , " &9 )+ &/ )O AQA I MfPq - ` (! " O E ," > )+ &9 )+ &/ % 17 b= ` )9 ? * aI5d ,' = L/ B ,> c % !" # $ ,' "? : c ? O !F = & > 1( U = / 1= ? ? )9 RdI^_ JQ ? > ! 1... ’ ? F c M g• 5_ T 5R M g c 5_• • dd • ^1B M • 5dd T dI dd qs H B = ,% # = )9 ! T J 7H? " XdI dd] _? > , = 14 e Z • d 5_ &4 /? ` `& fQB ! E / |... R_T ?bŠ X Xs & 55 Rd ` ? )8 ? ? d 5_T ?bŠ W Xs & ? G ! ` ` i ; ^u W 5u R^ ? ! ` T Jx ? 14 [ G YdI Rd ] _ ? ? ` : ," P` , nn` l s AQA l s R 5_ d ^d 5_ d F & )K )+ `, Rdu Q` `1 1‹H l s †? ` K