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Reproductive Biology and Seed Production of the Tropical Abalone Haliotis varia Linnaeus (Gastropoda) THESIS SUBMITTED IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE DEGREE OF 'Doctor of Plii£osopfty IN MARICULTURE OF THE CENTRAL INSTITUTE OF FISHERIES EDUCA TION (DEEMED UNIVERSITY) VERSOV A, MUMBAI - 400 061 'By T.M NAJMUDEEN CENTRAL MARINE FISHERIES RESEARCH INSTITUTE (INDIAN COUNCil OF AGRICULTURAL RESEARCH) P.B NO: 1603, KOCHI- 682014 INDIA SEPTEMBER 2000 Reproductive Biology and Seed Production of the Tropical Abalone Haliotis varia Llnnaeus (Gastropoda) THESIS SUBMITTED IN PARTIAL FULFILMENT OF THE 'REQUIREMENTS FOR THE DEGREE OF tIJoctor o!Pfii[osopfiy IN MARICUL TURE OF THE CENTRAL INSTITUTE OF FISHERIES EDUCATION (DEEMED UNIVERSITY) VERSOV A, MUMBAI - 400 061 13y T.M NAJMUDEEN CENTRAL MARINE FISHERIES RESEARCH INSTITUTE (INDIAN COUNCIL OF AGRICULTURAL RESEARCH) P.B NO: 1603, KOCHI- 682014 INDIA SEPTEMBER 2000 CENTRAL MARINE FISHERIES RESEARCH INSTITUTE (INDIAN COUNCIL OF AGRICULTURAL RESEARCH) \lTC¥~ ICAR Dr A.C.C Victor Tuticorin Research Centre of CMFRI, Tuticorin-l Senior Scientist CERTIFICATE This is to certify that the thesis entitled "Reproductive biology and seed production of the tropical abalone Haliotis varia Linnaeus (Gastropoda)" is an authentic record of the research work carried out by Mr T.M NAJMUDEEN under my scientific supervision and guidance at Central Marine Fisheries Research Institute, Kochi, in partial fulfilment of the requirements for the degree of Doctor of Philosophyof the Central Institute of Fisheries Education (Deemed University), Mumbai, and no part thereof has been presented for the award of any other degree, diploma or associateship in any University ~ Kochi - 682 014 September, 2000 DR A.C.C VICTOR (Supervising Guide) DECLARATION I, T.M NAJMUDEEN, hereby declare that this thesis entitled Reproductive biology and seed production of the tropical abalone Haliotis varia Linnaeus (Gastropoda) is a genuine record of the research work done by me under the scientific supervision of Dr A.C.C VICTOR, Senior Scientist, TRC of CMFRI, Tuticorin, and has not previously formed the basis for the award of any degree, diploma or associateship in any university Kochi - 682 014 September, 2000 I \\))~" , -I~ _,,-' T.M NAJMUDEEN ADVISORY COMMITTEE Chairman & Major Advisor: Dr A.C.C Victor Senior Scientist Tuticorin Research Centre of Central Marine Fisheries Research Institute Tuticorin-l Co- Chairman Dr P.C Thomas ~ (Y'-fl ~ Senior Scientist Central Marine Fisheries Research Institute Kochi-14 Members Dr K.K Appukkuttan Head, Molluscan Fisheries Divison Central Marine Fisheries Research Institute, Kochi- 14 Mr D Kandasami Senior Scientist Madras Research Centre of Central Marine Fisheries Research Institute Chennai- 06 Mr K Narayana Kurup Senior Scientist Central Marine Fisheries Research Institute Kochi-14 ACKNOWLEDGEMENTS I liadgreat privilege to wor/t under tlie a6fe guid'ance of(/)r }l C C Victor, Senior Scientist, 'l'1?.C of C9IffF(jlj, q'uticorin I ta~ tliis opportunity to express my deep sense of gratitude for suggesti11{J tFiis pr06fem andfor givi11{J me inva{ua6fe guid'ance witFi constant encouragement I am gratefu{ to my advisory committee mem6ers, (/)r CP C rzTiomas, Senior Scientist, 9rlr 1( Narayana /turup, Senior Scientist, (])r 1( 1( )lppu/t/tuttan, Jfead, ::MP(/), C::MPCJ?j, 1(oclii and::Mr (j) 1(andasami, Senior Scientist, ::M1?.C of C::M P(j{I, ::Madras for tlieir sincere lie[p offered me during tlie course of my wor~ Witli mucli gratitude I tlian( (])r 0/ 1(ripa, Senior Scientist, ::MP(/), C9rlPCJ?j, 1(ocFii, for critica{{y correcting tFie manuscript ana for providing va{ua6fe suggestions 7Fie guid'ance of (/)r Jl qopafa/trisFinan, Senior Scientist, mpq~ CocFiin Ce~ is a[so gratefu{{y ac/tnowfedfJetf I am FiigFi(y inde6ted to 9rlr {}306y Ignatius, Scientist, ::Mr 9rl {}3adarutfeen, former rr'eclinica{ Officer and ::Ms Preetlia Pani/(/tar, Scientist, 9rl1?.C of C::MPCJ?j, ::Matufapam, for tFieir unfaiung cooperation and advice tFirougFiout tFie period of my wor~ It is my privifege to express my Fieartfe{t tlianR.§ to 9rlr :JV.1( Sana~ Scientist, (])r 1( CR,gngarajan, former Senior S~ntist, Mr 9rl jIyyappan Pilfai, q'ecFinica{ Officer atuf 9rlr 1?g.gFia'Van, PFiotograpFier, C9rlP(j(j, 1(ocFii, for tFieir Fie(p in carryi11{J out f£fectron microscopic studies lowe tlie deepest sense ofgratitude to (j)r Pau{ (j{aj, Officer -in-cliarge, atuf(j)r C Susee{an, former Officer-in-Cliarge, Pq(]!rofessor, (])epartment of Zoo{ogy, :Mr (])hannaraj, Senior Scientists, rtrJ