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/H /.¥ ILLUSTEATIONS OF NEW SPECIES OF EXOTIC BUTTERFLIES BY WILLIAM C HEWITSON SELECTED CHIEFLY FROM THE COLLECTIONS OF W WILSON SAUNDERS AND WILLIAM VOL C HEWITSON IV JOHN VAN VOORST, LONDON 1S67— 1871 \c\o^A-3 LONDON : PRINTED BY WOODFALL AND KINDER, MILFORD LANE, STRAND, W.C iME ' M H O R P I D M CLEROME CLEROME BESA Upperside Eemale, rufous or yellow-brown, darker towards the margins Underside brown Anterior wing with two cell, an oblique short line white Posterior wing crossed dots near the apex before the middle white and iris is lines within the bands of brown linear ocelli, and between them three minute white each with the pupil dots Hab Borneo in In the Collection of This species brown by two beyond the middle two large black ; rufous, Expan 2^7 indistinct from the middle of the costal margin and three minute W C Hewitson closely allied to C Phaou of Erichson, but differs from it in wanting most of the transverse linear bands of the underside in having the ocelli of different form and colour, and three minute white dots between them CLEROME CHITONE Upperside Female, rufous-orange 2, the apex broadly : brown with some of the decorations of the underside seen through Underside paler Anterior wing with a black spot within the an irregular band of black the middle, and two beyond the middle : linear with a spot at the base, a transverse line near costal margin zig-zag) all the iris (indistinctly traced black ; it, : ocelli, in In the Collections of Saunders and Posterior wing lines (the inner line each with white pupil and ray of blue dots, Hab Celebes W W at an oblique broad white band beyond) and two submarginal two large black band by a short line from the middle of the yellow margined with black Expan 3xjj inclosed the disco-cellular nervures, a linear transverse submarginal bands black an ocellus and minute white spot near the apex : cell W C Hewitson MORPHIDjE CLEROME CLEROME MEN ADO Upperside Male (fig 4, orange-brown, slightly darker towards 5), the outer margins Underside brown costal Anterior wing crossed obliquely from the middle of the margin by a narrow band of white end of the a transverse curved cell, a spot and two lines within the : band Posterior wing with a spot margin, the black all ; two large black ocelli • three minute white two transverse undulated at the base, bands before the middle, two submarginal bands the middle touching the white band, at the two undulated submarginal bands, and the outer margin, black dots near the apex cell, (the inner band zig-zag) each with white pupil and ray of and the lilac dots : yellow, margined with black iris Female underside ; differs little from the male, except in colour ; it is much the bands also which cross the anterior wing approximate paler on the more closely at the inner margin Expan Hab Menado In the Collections of W W Near of The little in to C Stomphax butterfly, resemblance Saunders and W which Mr Westwood has placed to the other species He has, it is of its divergencies, but, besides these, it has others longer Would distance from it C Hewitson Westwood ; in this true, put genus under the name of C Busiris, has it into a subgenus, the eyes are not be better placed with the Nymphalidse, and if and pointed out some more prominent, and the body much not in the genus Iaera at no great it'.' DSI • iewitscm, del I eT: ± HYANTIS FricV \: DRUSILL 3.4 DRUSILLA BIOCULATUS 5.6 HYATT TIS HODEVA Tjv D1MONA : M O R P DRUSILLA H & I D HYANTIS DRUSILLA BIOCULATUS Morpho I 1, Hyades Indra bioculatus Guerin, Voy Coquille } Pl 17 Boisd., Toy Astrolabe, p 158 Upperside Female, white ; anterior wing rufous-white the costal margin, the apex, and the outer margin to a little beyond the middle (covering one-third of the wing), dark brown the costal and outer margins dark brown Posterior wing orange-yellow a large dark brown : ; : middle of the costal margin, inclosing two black ocelli with the pupils and the ocellus nearest the inner margin has a third minute ocellus attached to it and circular spot near the iris light blue ; inclosed within the same iris Underside as above, except that the costal margin of the posterior wing is of a much darker brown and is marked beyond the middle by an ocellus similar to the others, but smaller that the central spot is much larger, extending to the inner margin, and of a darker brown that the ocelli are much larger, especially that nearest the inner margin which has absorbed the : small ocellus mentioned above and contains two pupils Expan 3^ in In the Collection Hab Waigiou of A R Wallace It was not until after the accompanying plate was finished, and I believed that I had figured a new species, when I ascertained that it is identical with the Morpho bioculatus of Guerin It is not, as supposed by Boisduval and Westwood, the female of Melanitis (Dyctis) Agondas, from which it differs in the position of the nervures It is a Drusilla, and was placed in that genus by Dr Boisduval under the name of Hyades Indra The Dyctis bioculatus of the " Genera of Diurnal Lepidoptera," Plate LIV.* is not the Morpho bioculatus of Guerin but the true female of Agondas and not being able to discover any resemblance to the butterflies then before me in the figure given of Dyctis Agondas in the " Voyage de TAstroIabe," I figured it and a series of the females in the "Proceedings of the Zoological Society," Annulosa, Plate LV under the name of Melanitis Melane D bioculatus, though agreeing with the other characters of Drusilla, differs from all the other species or varieties in the arrangement of the discocellular nervures of the anterior wing in D Horsfieldii the first and seeond disco-cellular nervures are very short and of equal length, the third very long and curved; in D bioculatus the first disco-cellular nervure is very short, the second very long, and the third short On examining the numerous examples of Drusilla in my collection, I find that there is considerable difference in the length of the second disco-cellular nervure iu Horsfieldii, as stated above, the first and second disco-cellulars are of equal length in all the others the second nervure is three or four times the length of the first ; ; ; : DRUSILLA DIMONA 3,4 Upperside Male, white; anterior wing with the costal margin and apex broadly rufousthe outer half dark brown, with two unequal light blue near the outer margin, each pupilled with light blue, the larger irrorated with black brown ocelli : posterior Underside large ocellus is wing rufous-white as above, except that the base of the posterior all black with an and again encircled by apex with the blue Expan 3^ : in iris a iris of blue common band : wing is dark brown : that the that both of the ocelli are bordered with Mark, of orange ; and that there is a third ocellus near the incomplete Hab Aru W YV Saunders and W C Hewitson stated my belief, and I may repeat it here In the Collections of I have elsewhere (excepting D Horsfieldii from the list in consequence of its differing in the length of the disco-cellular nervures) that D Catops and Selene of Boisduval, D Phorcas and Milsecha of Westwood, D Myops and Macrops of Felder, and D Artemis, Anableps, and Dioptica of Volleuhoven, are only varieties of Urania D Dimona should, very probably, add one more to the number it differs, however, so much from all ou the upperside, that I have preferred for the present to cousider it as distinct ; ;; DRUSILLA AND HYANTIS MORrHIDiE HYANTIS Hewitson Body small, quadrate, abdomen slender Eyes small, smooth Palpi rather long, comAntennae rather short, thickened pressed, clothed with scales throughout, encircling the eyes gradually to the point, the articulations numerous and easily seen Anterior wing rather large the costal margin arched, apex rounded, outer margin slightly sinuated at the middle, inner margin rounded, protruded as in Euploea ; costal nervure swollen two at its base, reaching beyond the middle of the wing subcostal nervure with four branches together just before the end of the discoidal cell, the third midway between the end of the cell and the apex of the wing, the fourth at one-third of the distance between the last and the apex upper disco-cellular nervure very short, discoidal cell nearly half the length of the wing, closed : : : : the second semi-circular arched inwards rather short, the third more than twice the length of the second curved united to the third branch of the median nervure at a distance from its base wing oblong, circular, with large abdominal fold costal nervure joining the margin subcostal nervure with its first branch arising near the base considerably beyond the middle the upper disco-cellular nervure long, of the wing the discoidal cell short and narrow, closed Posterior ; : ; : branch of the subcostal nervure, at first in a transverse longitudinally down the wing the second half its length, direction, and afterwards passing its beyond base little joining the third median nervure a arising at a short distance below the first : Fore legs of both sexes short, and tibiae both so densely clothed with hair as tarsi of to conceal the joints of the tarsi Hind smooth legs long, slender, : the femora tibiae and tarsi of equal length HYANTIS HODEVA 5, both wings with the costal margin and apex broadly brown, the beyond its middle anterior wing rounded and produced at its inner margin as in Euploea near this margin a large oval pale rufous spot near the apex a Posterior wing, with a small spot of white and a black ocellus with some minute white dots small black ocellus with white pupil, broadly bordered with yellow, and marked by a very indistinct circle round the ocellus Upperside Male, white outer margin also brown ; to ; : : Underside as above, except that the ocellus at the apex of the anterior wing has the pupil and some smaller dots of white and an iris of yellow that the ocellus of the posterior wing that there is larger and has several minute dots of white and the iris bordered with brown is a similar ocellus on the costal margin bipupillated and clotted with white : : Female larger the wings broader, suffused with darker brown, the nervures dark brown wing with the centre alone white posterior wing white only to the middle, tinted with orange-yellow towards the inner margin the ocellus small, indistinctly pupilled with white the iris orange-yellow, bordered with brown and encircled above by a pale hue underside it differs only in the broader outer margin of the posterior wing : anterior ; ; : ; Expan S 3^% ? jq Hab Waigiou In the Collection of A E Wallace This is a species of singular interest, from the remarkable resemblance it bears to the Drusillas, differing at the same time from them widely in the arrangement of the branches from the subcostal nervure of the anterior wing, in the form of that wing with its projected inner margin, and in the closed discoidal cell of the posterior wing It is unique in the collection of Mr Wallace EMrcram EURY&ON/L VII | EURYGONA CLE (NAEUROMUS - , A EUBA&] F ' i E RYC I N I D M EURYGONA VII EURYGONA EUGEON 62 Male dark brown The apex of the anterior wing lighter brown, tinted with lilac Both wings crossed transversely by two bands of brown and a row of small indistinct spots near the outer margin (scarcely seen on the anterior wing) Eemale differs only in being of a much lighter brown Expan 1-^q in Hab Amazon In the Collections of W W Saunders and W C Hewitson Upperside Underside lighter EURYGONA EURYPUS 63 Male dark brown Both wings crossed transversely, at the middle, by a band of red-brown Posterior wing crossed beyond the middle by a second band of the same colour Both wings with some small indistinct spots near the outer margin Expan ly^- in Hab Amazon Upperside Underside lilac-brown In the Collection of Mr Bates EURYGONA EUBOTES 64 Upperside Male dark brown The anterior wing, and the posterior wing its costal margin to the middle, glossed with blue Underside light brown tinted with lilac Both wings crossed by a transverse Posterior wing with a subline of red-brown (broken where the wings meet) marginal row of small triangular spots (white at the base, black at the point), the from middle spot larger than the rest Expan l-2no Hab Amazon In the Collections of W W Saunders and W C Hewitson ERYCINID.E — EURYGONA EURYGONA EUHEMERUS Ewygona Euhemerus Doubleday, British 65, 66 Museum Catalogue Female sometimes of a uniform brown, sometimes with a large on the anterior wing Underside rufons Both wings crossed at the middle by a continuous curved Posterior wing with a submarginal row of small white spots, of red-brown Upperside central spot of orange line touched with black Expan l-gij Hab Amazon In the Collection of W W Saunders EURYGONA E Clem Upperside Underside continuous baud Expan l^f CLESA Male dark brown, glossed with of MS Boisduval rufous, tinted with lilac ; 67 blue both wings crossed at the middle by a red-brown (broad on the anterior wing) in Hab Brazil In the Collection of Dr Boisduval EURYGONA EUBAGES 68, 69 Upperside Female, sometimes of a uniform light brown sometimes crossed near the middle by a broad, ill-defined band of white, extending from the centre of the anterior wing to the inner margin of the posterior wing ; Underside lighter brown ; the transverse band commencing at the costal margin of the anterior wing Both wings with, near the outer margin, a row of lunular brown spots bordered with white, posterior wing with the middle spot large and black Expan 1-j^j in Hab Amazon In the Collection of Mr Bates EURYGONA EUROMUS Upperside Male black, with a large space of dark blue from the inner margin to the centre of both wings Underside The fringe white rufous, glossed with by a continuous waved 70, 71 line of lilac red-brown Both wings crossed, beyond the middle, Posterior wing with three or four small indistinct spots of white near the anal angle Expan 1^ in Hab Amazon W W Saunders surprise me if this should In the Collection of It would not prove to be the male of E Euhemerus eurygora vin W.CJIewitsan.dcl et litk prmtelLjHullmimtU 8cWJtun 1859 72 73 74.75.76 77 78 EURYGONA ZARA EURYG-ONA ZENA EURYGONA MAZACA 79 80 81 ? EURYGORA GYDA EURYGGNA DORINA ; ERYCINIDJE EURYGONA EURYGONA ZARA VIII $ 72, 73 E Zara, Hewitson in Gen Blur Lep., $ PL 71, fy Upperside Male dark brown, glossed with blue from the base to beyond the middle the outer half of the posterior wing dark blue the margin spotted with ; ; white Both wings light grey-brown, surrounded by a broad band of outwardly by bands of black and white alternately, three of each followed orange, anterior wing by a band of orange divided on the posterior midway on the separated Underside ; wing by white nervures Female (figs 72, 73) dark brown, lighter towards the base, tinted with purple on the posterior wing Anterior wing crossed beyond the middle by a large oval Underside as in the male, except that it has the white spot as above, white spot and less of the orange band of the anterior wing Expan In) Hab W C in In the Collection of Brazil Hewitson EURYGONA ZENA Upperside carmine 74, 75, 76 Male black Anterior wing, except the costal and outer margins, wing with a carmine spot near the middle of the outer margin Posterior Underside grey-brown from the base to the middle beyond the middle black, crossed by two bands of white the outer band of the posterior wing broken in two ; ; and projected towards the outer margin two orange spots near the anal angle Anterior wing with the base of the costal margin white, and a submarginal line of places ; orange towards the anal angle Female lighter brown, with a large spot of orange on the centre of the anterior wing, and one of the same size touching the outer margin of the posterior wing Underside from the base to the middle, light yellow; beyond it In-own, crossed, as in The outer margin of both wings orange towards the male, by two bands of white the anal angle Expan * l 2- > Hab Amazon Sla'oi 11 W W Saunders and W C Hewitson Iu the Collections of ?St - ERYCINID.E EURYGONA EURYGONA MAZACA 77, 78 Upperside Male brick-red, the margins broadly black except the inner margin wing and part of the outer margin of the posterior wing of the anterior Both wings crossed near the middle by a linear band with black line and white spot at the anal angle Posterior wing a of black Anterior of lunular white spots bordered with black, one spot row wing with a submarginal black above The outer margin towards the anal an^le near the middle broadly Underside grey-brown orange Expan l/o in In the Collections of Hab Amazon W W Saunders and W C Hewitson EURYGONA GYDA 79 Upperside rnfous-yellow Anterior wing with the costal and outer margins Posterior wing darker broadly brown, and a small black spot at the end of the cell on the outer margin Underside - Expan l of a uniform silvery white in In the Collection of Hab W Brazil C Hewitson EURYGONA DORINA 80, 81 The margins black A small black spot common wing with the costal and outer margins broadly Anterior to the base margin black with the outer wing Posterior black Upperside Male carmine of both, wings Both wings crossed beyond the middle by a darker rnfons Posterior wing with a small black and white spot (black above, white below,) Underside rufons band near the middle of the outer margin Expan H^ in In the Collections of Hab Amazon W W Saunders and W C Hewitson ' ; E EOT GIN IBJE EURYGONA IX w \ / si V V ' > s • ; ^0 v W C.Hewitson 82 AsA otliftv April KStNHa.r,ksu-t 15*1870 EURYGONA ERYTHRAEA 85 EURYGONA BETT1NA EURYGONA EFFIMA 82.83 84 8S 87 89 30 EURYGONA ONORATA EURYGONA PRAE CLARA .EURYGONA LABIENA 32 EURYGONA ATHENA - 31 am ERYCINIDiE EURYGONA IX EURYGONA ERYTHRAEA 82, 83 Eurygona Erythraea Bates Cat of Erycinida,p 375 Linn Soc Journal Zoology, Vol IX Upperside Male, scarlet Anterior wing with the costal and outer margins dark brown Posterior wing dark brown, with a large scarlet spot from the middle to the outer margin Underside grey-brown from the base to the middle dark brown from the middle to Posterior wing with the subnear the outer margin, traversed by three bands of white submarginal a line of orange at the marginal white band curved outwards at the middle ; : anal angle Exp \^q inch In the Collections of Hab Ecuador W W Saunders (Buckley.) W C Hewitson EURYGONA EFEIMA and Eurygona Effima, Hewitson Upperside 84 Equal Lep p 46 Male, dark brown, with the anal angle of the posterior wing broadly white Underside bands of brown by a black with white Exp spot : brown the outer margins brown Both wings crossed by three Anterior wing with a submarginal band of brown marked at its middle Posterior wing with a submarginal series of black spots bordered below pale : the third spot from the apex larger than the rest Hab Ecuador (Villano, Buckley.) 1^ inch In the Collection of W C Hewitson Does not differ from E Euryone, except in the white appearance of a small specimen of E Phedica EURYGONA at the anal angle BETTINA Eurygona Beltina, Hewitson On the upperside it has the 85 Equal Lep p 46 Upperside Male, dark brown, paler towards the apex of the anterior wing Underside glossy golden white Posterior wing with a single minute black spot at the middle of the outer margin Hab Ecuador W C Hewitson Exp l-j% inch (Angus, Buckley.) In the Collection of Altogether like E Eusepus of Hewitson, except that has a golden tint instead of the silvery white of Eusepus Mr Belt has sent me this species from Nicaragua it is EURYGONA ONORATA Eurygona Onorata, Hewitson without the band of the underside, and 86, 87 Equal Lep p 47 the costal Upperside Male, rufous Anterior wing with the centre orange margin a quadrate spot at the end of the cell, which is darker than the rest, the apex D D : Published October 1st, 1870.] EURYGONA ERYCINID.E and outer margin dark brown Posterior wing with the outer margin from the apex to the middle brown Underside Anterior wing grey-brown, with three central white spots Posterior wing white, with the base, a band from the costal margin to the middle, a curved line below this, and the apex brown a black spot bordered with white near the middle of the outer margin Exp Iyq inch Hab Ecuador (Sarayaco, Buckley.) : EURYGONA LABIENA Ent Mon Eurggona Labiena, Heicitson SS Mag p 226, Vol VI., 1870 Upperside Male, orange Anterior wing with the costal and outer margins and Posterior wing with the costal margin, which is broad, and apex broadly dark brown the outer margin, which is narrow, dark brown the inner margin paler brown Underside orange-yellow Posterior wing with a submarginal series of minute black ; spots Eemale like the male, except that it is yellow instead of orange Exp l-g ^ inch Hab Nicaragua (Chontales, Belt.) In the Collection of W C Hewitson EURYGONA PR^ECLARA Eurggona Prceclara, Hewitson is 89, 90 Equat lep p 48 Upperside Male, orange Anterior wing with the costal margin and apex (which broad) dark brown Posterior wing with the apex brown Underside Both wings crossed beyond the middle by a series of pale grey spots, and by a submarginal series of minute brown spots Anterior wing crossed at the middle by a band of four rufous spots Posterior wing with a spot near the base, three spots at the middle, and three spots near the anal angle, all rufous Exp 1-j^ inch Hab Ecuador (Sarayaco, Buckley.) In the Collection of W C Hewitson Very near having the apex more pointed and broadly bordered head from the light When reversed it is one of the most glorious things in this part of the Creation, surpassing the Morphos wit!) brown I to E Opalina, from which it have described it as seen with differs in its EURYGONA ATHENA Eiirygona Athena, Hewitson 91, 92 Equat Lep p 46 Upperside Eemale, dark rufous-brown the outer half of the posterior wing (which is rounded at the outer margin) orange Underside rufous-grey Both wings crossed at the middle by a rufous-orange band, paler on the posterior wing and bordered on both sides with brown both wings crossed beyond the middle by a less-defined rufous band, which ceases at the middle of the posterior wing Anterior wing darker towards the outer margin Posterior wing with its outer half orange-yellow the outer margin orange a submarginal baud of eight white spots, with a rufous border on both sides, except the middle spot (which is crowned by a black spot) and the three spots nearest the anal angle (which are bordered above and below with black) Exp l^- inch Hab Ecuador (St Ines, Buckley.) In the Collection of W C Hewitson : : : : E URY&ONA.X 1UU V / WW 10* "W C Hmtson del etlith JuVl " 18 72 M.fe T) Ha-r"ka;rt imp EURYGOEJA EURYSTHENES 95 9G 97 EURYGONA EUCERUS 101 102 EURYGONA EUCRATES 103 104 EURYGONA EURYMACHUS 98 99.1OO.EURYG0NA ARGEETTEA 93.94 - _ ERYCINID^ EURYGONA EURYGONA EURYSTHENES Upperside 93, 94 Both wings bordered at the broad on the anterior wing Male, black brilliant lustrous blue, very outer margin with Anterior wing crossed at and beyond the middle Posterior wing crossed at margin by straight rufous bands the middle by one baud of the same colour followed near the outer margin by one a large lustrous blue spot, bordered with orange except at its base, which is white submarginal series of black spots and above them hastate orange spots bordered with brown the outer margin rufous Underside and lilac-rufous at the outer : : This species and E Eurymaehus are When reversed the brilliant blue Exp l-j^- makes its Hab Ecuador inch W Iu the Collection of totally black when their heads turned to the light appearance (Buckley.) C Hewitson EURYGONA EUCERUS 93, 96, 97 Male, dark brown with a large spot from the base to the middle Posterior wing rufous with a large orange spot rufous at its base, orange beyond near the costal margin which is dark brown Posterior Anterior wing pale-brown rufous near the middle Underside at the middle crossed with base wings brown at the Both wing lilac-white clouded brown spots, and some spots, beyond the by middle by a broken band of red-brown with on the white spots, above black bordered near the outer margin by a series of Upperside anterior wing Exp ; % l-2 the central spot of the posterior wing largest inch Hab Brazil (Rogers.) In the Collections of W W Saunders and W I have figured 97 as a variety of this species C Hewitson It does not differ on the upperside EURYGONA ARGENTEA Eurygona Argentea, Hewitson 98, 99, 100 Trans Ent Soc p 166, 1871 brown Anterior wing rufous below the median the middle of the wing, where it ends in a more distinct round spot Posterior wing with a longitudinal rufous-orange band from the base to the outer margin Underside silvery white Anterior wing with four transverse bands and the outer margin brown Posterior wing crossed by six brown bands, all tending Upperside Male, dark nervure, from the base towards Published 1st July, 1S72.J ERYCINID.E EURYGONA towards the anal angle two from near the base, a third from the costal margin united which runs near the inner margin, the other two near the outer margin, the outer margin black, marked by a the outer one marked by a small black spot large orange spot Female like the male except that there is on the upperside a large orange-yellow spot on the anterior wing, a small undefined rufous spot at the middle of the posterior wing, and a small spot of the same colour on the outer margin near the anal angle : to the fourth ; The spot on the posterior wing Exp - inch is so rubbed off that I have Hab Nicaragua l-2 In the Collection of Thomas had to guess its form and extent (Chontales.) Belt EURYGONA EUCRATES 101, 10:2 Upperside Male, red-brown Anterior wing with the costal and outer marPosterior wing crossed longitudinally between gins and apex broadly dark brown brown band of dark nervures by a the median Underside ochreous-brown Bothwingswiththeba.se dark brown, both crossed beyond the middle by a linear brown band bordered outwardly near the costal margin by a large brown spot Posterior wing with the inner margin below its middle to A submarginal series of white spots crowned with the anal angle broadly white large and triangular spot from the apex third black, the (Buckley.) Ecuador 1-^ inch Hab Exp In the Collection of W, C Hewitson EURYGONA EURYMACHUS 103, 104 Anterior wing with a large spot of lustrous blue near Male, black the anal angle, posterior wing with a similar spot at the apex Underside brown Anterior wing crossed at the middle by a distinct band of dark red-brown, beyond the middle by an undefined broad band of brown and Uppekside Posterior wing the outer margin rufous submarginal band of brown by a large margin crossed at the middle by a rufous band followed near the outer black small series of dark blue spot bordered with orange its base white and a brown with bordered triangular spots with above them hastate orange spots broadly and below them white the outer margin rufous (Buckley.) Exp l-^o inch Hab Ecuador by a : : In the Collection of W C Hewitson ... ; two obscure iris C Hewitson This species, though much resembling T Camadeva of Westwood, the basal half of the wings is altogether different the middle of the wing are of another form ; : five... Boisduval under the name of Hyades Indra The Dyctis bioculatus of the " Genera of Diurnal Lepidoptera," Plate LIV.* is not the Morpho bioculatus of Guerin but the true female of Agondas and not being... name of Melanitis Melane D bioculatus, though agreeing with the other characters of Drusilla, differs from all the other species or varieties in the arrangement of the discocellular nervures of