12345678910. Thomisidae aus Japan. I. Das Genus Tmarus Simon (Arachnida: Araneae).Acta arachnol., 27 (spec. no.): 6184 (1977).. Thomisidae aus Japan. II. Das Genus Oxytate L.Koch 1878 (Arachnida:Araneae). Senckenb. biol., 58: 245251 (1978).. Thomisidae aus dem NepalHimalaya. I. Das Genus Xysticus C.L.Koch 1835(Arachnida: Araneae). Senckenb. biol., 59: 267288 (1978).. Thomisidae aus dem NepalHimalaya. II. Das Genus Lysiteles Simon 1895(Arachnida: Araneae). Senckenb. biol., 60: 91108 (1979).. Fossile Spinnen aus miozanen Sedimenten des Randecker Maars in SWDeutschland (Arachnida: Araneae). Jh. Ges. Naturkde. Wurttemberg,134: 133141 (1979). (W.Schawaller t 4E 1t). Thomisidae aus Japan. III. Das Genus Lysiteles Simon 1895 (Arachnida:Araneae). Senckenb. biol., 60: 203217 (1980)
Thomisidae aus Japan I Das Genus TmarusSimon (Arachnida: Araneae) Acta arachnol., 27 (spec no.): 61-84 (1977) Thomisidae aus Japan II Das Genus Oxytate L.Koch 1878 (Arachnida: Araneae) Senckenb biol., Thomisidae aus demNepal-Himalaya I Das Genus Xysticus C.L.Koch 1835 (Arachnida: Araneae) Senckenb biol., 59: 267-288 (1978) Thomisidae aus demNepal-Himalaya II Das Genus Lysiteles Simon 1895 (Arachnida: Araneae) 58: 245-251 (1978) Senckenb biol., 60: 91-108 (1979) Fossile Spinnen aus miozanen Sedimenten des Randecker Maars in SWDeutschland (Arachnida: Araneae) Jh Ges Naturkde Wurttemberg, 134: 133-141 (1979) Thomisidae aus Japan III Das Genus Lysiteles Simon 1895 (Arachnida: Araneae) (W.Schawaller t 4E1t) Senckenb biol., 60: 203-217 (1980) Thomisidae aus demNepal-Himalaya III Das Genus Stiphropus Gerstaecker 1873, mit Revision der asiatischen Arten (Arachnida: Araneae) Senckenb biol., Erstnachweis einer Krabbenspinne (Thomisidae) in dominikanischem Bernstein (Stuttgarter Stuttgart 61: 57-76 (1980) Bernsteinsammlung: Arachnida, Araneae) Beitr Naturk., B, (73): 1-13 (1981) Revision japanischer Spinnen I Synonymieeiniger Arten der Familien Theridiidae, Araneidae, Tetragnathidae and Agelenidae (Arachnida: Araneae) 10 Acta arachnol., 30: 1-7 (1981) Verwandtschaft von Tetrablemmaphulchoki Lehtinen 1981 (Araneae: Tetrablemmidae) Senckenb biol., (1) 62: 349-353 (1982) 11 Zodariidae • aus dem Nepal-Himalaya Walckenaer 1805 (Arachnida: I Neue Arten der Gattung Araneae) Senckenb biol., (Araneae, Thomisidae) Storena 63: 211- 217 (1983) Eine neue japanische 12 Synaema-Art Acta arachnol., 31: 59-63 (1983) 13 The Thomisidae • of Japan IV Boliscus a genus new to the Japanese (A), 10: 63-71 14 Revision • einiger Japan 15 The Thomisidae • natn of Japan 16 17 Sci Sci Araneae), natn Tokyo, Sci Mus., Mus., Tokyo, (Arachnida, Araneae) • Japan (Araneae: (31): 14-19 (1985) A new spider 1869, and its Simon, 1903 (Arachnida, Araneae) • Mus., Little-known 19 • Spiders Pap pres species areas Jpn Japanese Mus., spider, of the East China Sea Clubiona Clubionidae, Japan Sci natn Sci Clubionidae) species-group Bull natn and Thomisidae from (1986) Mus., Tokyo, (2) Thomisidae) Bull (Araneae, Gnaphosidae and the southern (1986) (D.-X Song k) zilla the Noto Peninsula Tokyo, Clubionidae) Tokyo, pp.19-25 of the genus Cupa (Araneae, Tokyo, (A), 12: 117-121 Mem natn 'tool., (Araneae, Kurosawa, of a new and peculiar of the families Bull 1900, aus Soc syst corticalis Tokyo, (A), 12: 25-29 (1986) Representative Sci Proc of the group of Clubiona A new Sino-Japanese new junior (A), 11: 91-97 (1985) Thomisidae) Ent aus (A), 11: 19-39 (1985) Eine neue Art der Gattung Misumenops F.O.Pickard-Cambridge, • from the coastal 20 Thomisidae V Monaeses Thorell, Mus., Tokyo, found in Japan 18 Bull 1891 (Arachnida, (1984) synonym, Mecostrabus natn fauna Arten der Familie Bull Thorell, part (19): of Ishikawa 167-174 Prefecture, (1986) 21 • Twonew species of the family Zodariidae from Saudi Arabia (Arachnida: Araneae) 22 • Fauna Saudi Arabia, 8: 6-11 (1986) (R.Jocque ti 1l n) A new Japanese castianeirine genus (Araneae, Clubionidae) with presumptive prototype of salticoid Tokyo, (A), 13: 13-19 (1987) (3) eyes Bull natn Sci Mus., A Revisional Study of the (Arachnida, Spider Family Araneae) Thomisidae of Japan By Hirotsuau Department of Zoology, National Ono Science Museum, Tokyo • Contents Introduction • Acknowledgement A Brief History Material and Type of the Study • in Japanese Spiders • • of the • Thomisid Species Recorded from Japan • Terminology • • • • Abbreviations Taxonomy: Family Subfamily Thomisidae Stephanopinae Genus Cupa Genus Phrynarachne Subfamily Dietinae Oxytate Genus Loxobates Genus Alcimochthes Pickard-Cambridge, Thorell, L Koch, 1878 Sundevall, Tmarus Simon, Genus Monaeses Genus Coriarachne Genus Bassaniana Genus Xysticus C.L Genus Oxyptila Simon, • 1871 • • • • • • • • Strand, • • • • • • 1885 • Koch, • • • • • 1870 • • • • • • 1835 1864 -i- 10 • 12 17 • 20 • 21 27 • • 1833 1928 • • 30 38 1869 Thorell, • 1875 Thorell, • • 1877 Simon, Genus 1869 1895 Thorell, Thomisinae 1833 1906 Simon, Genus Subfamily Strand, • • Sundevall, • • Methods Depository • • • • 50 Genus Lysiteles Genus Synaema Genus Takachihoa Ono, 1985 Genus Misumenops F.O Pickard-Cambridge, Genus Diaea Genus Heriaeus Simon, Genus Misumena Latreille, Genus Pistius Genus Runcinia Simon, Genus Thomisus Walckenaer, Subfamily Incertae Nomina Higher Thorell, Boliscus Doubtful List Simon, Bominae Genus Species Simon, Records, 1895 1864 the Taxa Classification Zoogeographical Simon, Simon, 1804 • • • • • • • • • • • • • • • • • • • • • • 289 • • • • • • • 295 • • • • • • 303 • • • • • 311 • • • • • • 319 • • • 332 • • • • • • • 1875 • 1805 1886 1891 Sedis Misplaced and Misidentified Described • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • Species • in this Phylogeny Paper • • Figures • •• • •• • • • • • • • • • • • • • • • 241 • • • 258 • • 264 • • • 351 • • • • 353 •• • • • • • • • • • • 359 361 365 • • •• • • • •• ^• •• • • • • • • • • ^• • 220 • 1875 Notes of 1900 1875 Thorell, and • • 1869 References Explanation • Nuda of • • • • • • • • • • • • • • • • •• • 370 • • 366 389 399 447 -ii- Introduction The purpose thomisid spiders morphology, in Japan regions Since Japan, 70 specific spent years for first to read not only from Japan of the world, first institutions because very in the sense Other than those the type study consists from various places thomisid (details are given were placed of of the original specimens countries old collections since -1- in a later identification or original of most species Institute in Europe, with curators I of I visited of many on loan general of the specimens in Japan philodromids became inevitable During my stay specimens, their at the Zoological and communicated from in wrong taxonomic since spiders spider names excluding been recognized studying as well, the were based on doubtful of Mainz in Germany museums for examining myself spiders described from Japan and her adjacent used in this was necessary Thomisidae some species Re-examination the University other parts country and many have never several it and subspecific from this naming; descriptions of comparative on the family However, some records positions described subject, but from all L Koch (1878c) have been reported inappropriate history sciences more than chapter) through to the thomisid has been made generally modern biological the natural zoogeography this referring and her adjacent is to present by investigating to pursue literature little study taxonomy and partly In order through of this and comparative personally 1966 and those collected material by G \ i!r, } :j.'1)167 ::.!'//e „\J\ l _ ,_.(i)ek , , ',._ )_ _.,.\ , :: ,\ ir- t:;::,1,,,, 4:„ 4„ _., i• „: :, :.i.: ,„ ., ,: 169 168 ~ yjs , „,/ lif i- :,: P.: I , -t,„\j, , , ,„ r::,., , ki,:,-.s.,,,,,,, /j/ :,, N \s 'Ait: „, ; ,_ „ •::: 2,,.,,,it/l'-'.* oso ( opotex, 5w,:r.' —/ I '''.' , \ „ :, A 91'''''''' d,/ `r 170't 171 [ 7:'pe -: .?: CINIIIILa- •-•.• .:1::: •::•,,, 'n.i i :.i.':: -: :: : ::•.: • - - : •' : :'.1'•::::'.:: ::'i.::::;•: —":', •:J :: ::: ::::::::::• 'i ;:.- • : •'-,F.: ::.:::1*.;, •• ':' ':') ; r : : •-.',.;•;.•:::' -ig7••' :' • .••••—. • ::::::*: :.:- , :::.•:.:::.' 77. 7.:.•••fi:—.:•:-,: : .•:•••:::: •:.:•: 7-.'‘.^.: j.;., „-, -z ' _, -.:2.::'•••• ._ _ _ _ _ _ _ _. - - :'':.:•:,::.::' • -:•::.:'-'1:::••• ::::;:;;;-•'•'•::::'•,:-'••• • '•-•'•:'•::•: ::.::::: :•:::-:: ••:• ' •:.-:-., :1: l •• , • ,;., -1,:-':.,;.:.::.*:: ! :**:- •:: "'.'f'.•::::: :, ,, N;.: ,::: , y.:.!;.t ::: ,a _ \, _ . - — _ • • • •••:-::•:%:.i." :r : :•• ' ;:' - '1•,-. ., ')I • / , ,/ —^ '.:‘, 173 L I t\ , il1% 1"- _ •.•• ,„ : .;.• N , - \ \ 11 'iii tee asok •-• ' _,,,‘,•• • • • \ N, • — 174 172 0) , Ii ) I_ :., , ,,, , ,„.:-,•;:.•;+•.„1 At-01 • , „• :., ••,=•Ir: \ • ' ':', _157) ;•.- -.- - • !:.:••: • • , :- -, , „„ -: :::.: , , , .! : :'.1-: \ • : 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:^•^^^••' ." s-.1 - - I i iie*200 1111% lit, , l, •I •, — - 1.Q)_. r-j' E.'.: .6e, 7;:.-`\_.— , \ 199• - -/ — ,r,\— 201 // ,„:1,:s ,_.1._ „ _ , ,a; ,./ A(7.- 204 - i- f -R,‘!." * P^ :: ^ — ';': — 1,:s4:.T, •• ^ , ^ ft nt ,=4.,_ ''''::: - • - 7-,• - •';','' '.j ; 'I-°:c•-' v'•.'., : c-••- '''-t-1°f, t `' ‘-1 t -., • \:is., 4, .„1 , , • _ _ ' z , - ^ • , , , ' „ 202 203 / , ,s iliv id :::-" \,,,,206208 O E Q ; ::: 1:.77.7 '2' : PC ' VV c \"4V (‘-'.f 2)- 1-17-)''':, .i.:ii,W,; ,: ,, ,.:- ,2;:.i.i.;, f.': • , psi P c 205209 7• ' o ° r 207 214211`—` ~i~1 ~~~~ ", _Vir fs / / \ Y ','(213• 212 ;%;, 210 MN~J LM y"c\-o gars AM v ~c 5rJ' \`-/JNt-o~J dryLM s\ U \ p \O NR SR high altitude MM SR/ NRLM low Southern NorthernHonshuHokkaido Ryukyus RyukyusShikoku Kyushu Fig 215 IqP io r b Xvsticus labradore tritil° i411.511N1 cp { Xvsticus i Ca •p North Pole c_ ~ albidus Xysticus deichmanni v o.0 q bO /d ~\ Fig 216 I ^,' Xvsticus Xvsticus daisetsuzanus rugosus 4; Cf7 111(1 Fig 217 h a (8 ••'' •moo Q Jr 4— • • i1 • • p a ~ nn ll A•Of Alfo •o • o• Ad Lysiteles O L maius L okumae • L miniatus Fig 218 coronatus O b mod• Fig 219 V-)) 00 // O a 4II ° •• ,0 • •a• ^ • r,4'it G7~' a 0gT • • •.• Tmarus rimosus D • a4°• T makiharai I1 Fig T• 220 T yaginumai o V-O 0OQ 41 + , _.(-)fej VY1 Gf21° a, i 0,,, ;ti4+ F.: 0b Ail, a I• Runcinia AL R Fig y 221 acuminata albostriata T V-1 OQ a u - I T 'O e :.7.';'V-op : •• o~o a,» A 0G I T o r D o • t**** Fig 222 A • Thomisus • T labefactus kitamurai T okinawensis ... Thomisidae aus Japan I Das Genus TmarusSimon (Arachnida: Araneae) Acta arachnol., 27 (spec no.): 61-84 (1977) Thomisidae aus Japan II Das Genus Oxytate L.Koch 1878 (Arachnida: Araneae) Senckenb... Tokyo, (Arachnida, Araneae) • Japan (Araneae: (31): 14-19 (1985) A new spider 1869, and its Simon, 1903 (Arachnida, Araneae) • Mus., Little-known 19 • Spiders Pap pres species areas Jpn Japanese... dem Nepal-Himalaya Walckenaer 1805 (Arachnida: I Neue Arten der Gattung Araneae) Senckenb biol., (Araneae, Thomisidae) Storena 63: 211- 217 (1983) Eine neue japanische 12 Synaema-Art Acta arachnol.,