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m II AUb ^'Vfl- 193/ -^ BULLETINS AMERICAN PALEONTOLOGY VOL XV January ig2g J^7 —J^^h' Jy Harris Co Ithaca, N Y 11 S A ^93'^^ CONTENTS OF VOLUME XV Number 55 A Lower Tertiary Fauna from Manta, Ecuador P>y J J iuxlloway and Mars,aref Morrey 56 Three New Claiborne Fossils By 57 ir S i - - - '^ 1-56 57-66 Cole Contributions to the Tertiary Paleontology of Peru Part 57a A Pages Plates Upper Eocene Mollusca and Brachiopcria By A A, Olsson 2, : 16 67 17 117-122 18-21 123-142 22-80 i^:-3i6 New Oligocene Brachiopod from Mexico 1 fS r>y \V S Cole 57b Some Small Foraminifera from the Meson Formation of Mexico By 58 11: S and Ruth Cole Gillespie Higher Fossil Faunas of the Upper Aileglieny Bv A'einiefl! E ( 'asler ^^,,.»-''-.|Hs,r„^,^^ BUI^IvETINS OF AMERICAN PAI.EONTOLOGY Vol 15 No 55 A LOWER TERTIARY FORAMINIFERAL FAUNA FROM MANTA, ECUADOR By J J Galloway and Margaret Morrey Columbia L^niversitv New York, N Y # I Jamiary j, 1929 Harris Co Ithaca, N.Y U S A INTRODUCTION described and figured in this paper were Manta, Ecuador, by Mr M N Bramlette, of the collected near South American Gulf Oil Company, which company has kindThe material ly given permission for the fauna to be published was collected from a silty shale at the surface, and all the spe- The Foraminifera cies are Many from the same sample of the more fragile spec- imens have been broken in collecting or washing The fauna is of special interest in being the third and largest occurrence of small Foraminifera to be described from South America One species was described by Bornemann, from the Tertiary of Rio de Janeiro^, and twenty-five species were listed and five species and one variety described by Berry from Peru.- There are included in this paper eighty-eight species and vaThere are ten new sperieties, belonging to thirty-nine genera It is remarkable, in view of the fact that cies and one variet}' so few small fossil Formanifera have been described from South America, that there are so few new species in the Ecuador fauna The identifications have been made with extreme care, and in all cases where identifications have been made the Ecuador speci- mens are almost identical with the published figures The specimens have been compared in all cases, excepting that of Quinangnsta quelocidina The descriptions (Philippi), with the original figures and international Rules of Zoological ture have been strictly adhered to Nomenclanames In cases where the or definitions of genera are not agreed upon by recent authorities, the synonymies and descriptions of the genera are included The classification is that The age of the fauna of the senior author is not known to us except by what the The age cannot be told by Foraminifera themselves indicate the percentage of like species which have been reported different horizons The from t}pe specimens of the species w4th which Ecuadorean species have been identified came from the following horizons: ii from the Eocene, 15 from the Oligocene, 12 the Bornemann, 153, pi in 1, Erman's Archiv Wiss Kunde Russland, figs vol 14, 1855, p 5-15 Berry, Eclogae Geol Helvetiae, vol 21, no 1, 1928, p 130, figs 1-6 ; Bulletin 55 from the Pliocene, 17 from the Phocene, from tlie Pleistocene, and 18 from the Recent On the basis of specialized forms, such as A'odosorclla caiiicraiii (Dervieux), the age is upper Eocene The fauna bears greatest resemblance in general to the following Eocene and Lower Oligocene of Hungary, described faunas : Upper Eocene of Texas, described by Cushman and Applin Lower Eocene of Texas, described by Mrs Plummer Upper Eocene of Mexico, described by Cole and Pliocene of Kar Nikobar, described by Schwager There are no well known guide fossils, such as the orbitoids and numdescribed by Hantken ; ; ; mulites On we judge the whole, Eocene the fauna to be L^pper in age SYSTEMATIC DESCRIPTIONS Family MILIOLIDAE d'Orbigny, 1846 Genus QUINQUELOCULINA d'Orbigny, 1826 Quinueloculina angusta (Philippi) Plate Triloculina angusta Philippi, Tertiar nordwest Deutsch., 1843, 1, fig 43, p pi 40 Quinqueloculina angusta Reuss, CI., vol 18, 1856, p 253, pi Test elongate, elliptical, Sitz 9, fig Ak Wiss Wien, Math.-Naturw k 90 one side smooth; aperture at Length, 0.5 (Upper Oligocene.) flat, bers well marked, round in section tooth 3, fig ; the other inflated sutures depressed the end 'of a short neck, mm ; breadth, 0.23 mm ; cham- ; surface round, without Abundant Genus SIGMOILINA Schlumberger, 1887 Sigmoilina celata (Costa) Plate 3, fig Spiroloculina celata Costa, Mem Accad Sci Napoli, vol 2, 1855 (1857), p 126, pi 1, fig 14; Atti Accad Pontiana, vol 7, fasc 2, 1856, no description, pi 26, fig (Pliocene.) Planispirina celata Brady, Rep Voy Challenger, Zool., vol 9, 1884 p 197 pi 8, figs 1-4 (Recent.) ' Test subelliptical unequally biconvex in ; side view, the ends chambers very little somewhat pointed; inflated, about five Galloway and Morrey, EcUxVDor Forams 9 on a side; sutures obscure; wall thin, covered with tine sand the interior is smooth aperture terminal, round, or closeii with sand grains Length, up to 0.93 mm Abundant grains ; The ; wall very thin and the wall of the is broken out or uncompleted last chamber either is our specimens, and the apertures in all are closed Family GLOBIGERINIDAE Carpenter, 18fi2 Genus GLOBIGERERINA d'Orblgny, 1826 Globigerina bulloides d'Orbigny Pate 3, fig Globigerina bulloides d'Orbigny, Ann Sci Nat., vol 7, 1826, p 277, no 1; Modeles, no 17 and no 76; Voy Amer Merid., vol 5, pt 5, 1839, p 37; in Barker, Webb and Bertholet, Hist Nat lies Canaries, vol 2, pt 2, Foram., 1839, p 132, pi 2, figs 1-3, 28 (Recent.) Test composed of about ten globular chambers, rapidh* enlarging, four in the last ly perforate ; Length, the umbilicus into whorl surface reticu'ate sutures deep ; ; ; wall thick, coarse- aperture an elongate 0.28 mm breadth ; slit opening mm 0.23 Common Globigerina compressa Plummer Plate 3, fig Globigerina compressa Plummer, Univ Texas, Bull 2644, 1926 (1927), (Eocene.) 135, pi 8, fig 11 a-c p Test rotaliform, unequall\' biconvex, the ventral side more convex than the dorsal periphery lobulate peripheral margin sutures subangular chambers inflated, six in the last whorl depressed, curved on both sides of the test wall hyaline, medium surface granulate, aperture on the ventral coarsely perforate side, at the base of the last chamber, extending from the umbili; ; ; ; ; ; cus nearly to the peripher\', with slight Diameter, 0.2 lip mm Rare Our specimens are not as compressed as the Texas form, but otherw^ise they are very similar Plate Globigerina conglomerata Schwager Globigerina conglomerata Schwager, Novara-Exped., 1866, p 255, pi 7, fig 113 19, pt 3, 1926, p 745, Theil, 3, fig vol 2, (Pliocene.) Globigerina bulloides var quadripartita vol Geol text fig Koch, Eclogae Geol Helvetiae, (Middle Tertiary.) 20 ; 10 Bulletin 55 10 chambers elongate dorso-ventrally, inflated, the last one smaller than the penultimate, and usually depressed, four in the last whorl, forming a restricted, rectanguTest globular, high spired lar, ; deep; sutures straight-sided umbilicus or vestibule; deep, wall coarseh' perforate and granulate; aperture opening at the bottom of the vestibule, not generally visible from the exterior mm Length, up to 0.7 Abundant This species differs from the Pleistocene G quadrilaterata Galloway and Wissler (Jour Pal., vol i, 1927, p 44) in the larger size, elongate chambers, higher spire, and deep vestibule Plummer Globigerina pseudobulloides Globig«^rina pseudo-bulloides (1927), p 133, pi 8, fig • Plate Plummer, Univ (Eocene.) Texas Bull fig 3, 1926, 2644, Test oval in dorsal view, consisting of about nine inflated five chambers in the last whorl; spire depressed; su- chambers, tures deep ; wall coarsely perforate and granulate ; aperture like, extending from the umbilicus toward the periphery 0.33 mm ; breadth, 0.27 This species is mm slit- Length, Abundant related to G cretacea rather than to G bulloides from G cretacea in not having the aperture in die umand from G bulloides in having more chambers in a whorl, the chambers enlarge less rapidly, and the aperture is not It differs bilicus, in the umbilicus , Plate Globigerina trilocularis d'Orbigny Globigerina trilocularis d'Orbigny, Ann Sci Nat., vol 7, 1826, Fornasini, Kendi Sess R Accad Sci Isti Bologna, n — 1898, p 12, fig 2, pi 1, figs 6-7 3, fig p 277, no vol s., 2, (Pliocene.) Test composed of about seven or eight globular chambers, three or three and a half in the last whorl, the last nearly as large as the remainder of the test sutures deep ; ; wall coarsely perforate and granulate an arch opening into the umbilicus chamber spire depressed ; aperture Length, up to 0.4 mm Common This species differs from G triloba Reuss cessory dorsal apertures It differs in not having ac- from G bulloides only three chambers in the last whorl in having 311 Bulletin 58 172 Explanation of Plate 57 (78) FIGURE Ctenacanthus harrisi Jewitt, Pa Nat si::c sp From the Oswayo sandstone east of Mt approx P 103 n HoSoptychus tayiori (Hall)? From the Chemung- beds at Bradford, Pa Greatest width 38 mm P 104 Rhadinichthys sp nov East From the Dev Fishes of N Y., pi 19, Eothriolepis minor Newb fig P P 105 From Mon U S G S., No 16, pi 20, fig 106 Holoptychus americanus Leidy Ibid., pi., 19, fig 12, Rhadinichthys, sp nov East (See fig 3), pi 4, fig P 104 11 P 105 PL 78, Vol., 15 Hull Amer I'aleont No ,5s I'l 57 PL 58, No 58, P.UTviv Amur Pai.eont Vou 15, Pl 79 A^ 1' u ;?"l' I;, '^'*iwr^^' 12 '3 i5 61- H 16 ^^l» Caster: DEvoxo-CAKiioxiKEuous 315 Explanation ok Plate 58 17;> (79) FIGURE Dipterus (Ctenodus) fig nelsoni Newb Mon U G S S., No Dipterus Ctenodus Icvis Newb Ibid., pi., 27, fig 22 P 105 DipttTus (Ctenodus) levis Newb Ibid., pi 27, fig 22 P 105 Homacanthus acinaciformis pi 27 East From Dev Fishes of N Y., pi 1, fig pi 18, 106 16 P ' Apedcdi:." priscus Le;( y Ibid., pi Gyracanthus sherwoodi Newb From Mon U fig 16, 105 19 P P 1, fig 1, P 106 S G S., No 16, 10(i 13 Gyracanthus sherwoodi Newb Ibid., fig 4a Gyracanthus sherwoodi Newb From Dev Fishes of N Y pi fig Dipterus (Ctenodus) levis Newb Mon U S G S No 16, pi 27, fig 23 Dipterus (Ctenodus) ftabelliformis Newb Ibid., pi 27, fig 21 P 105 Dipterus (Ctenodus) nelsoni Newb Ibid., fig 20 P 105 Sagenodus sp From Dev Fishes of N Y., pi 1, fig 13 P 106 Dipterus (Ctenodus) quadratus Newb Mon U S.' G S., No 16, pi 27, 14 Dipterus (Ctenodus) quadratus Newb 15 Dipterus (Ctenodus) minutus Newb 16 Dipterus (Ctenodus) flabeiljformis Newb 10 11 12 fig 24 P 105 Ibid., fig 25 Ibid., pi 27, fig 26 P Ibid., fig 21a 105 Bulletin 58 174 316 Explanation of Plate 59 (80) FIGURE Spirodomus insignis Beecher From 39th Rep N Y pi 12, fig S Mus Nat Hist P 82 Spirodomus insignis Beecher Tropidocaris interrupta Beecher Ibid., fig P 97 From Pal of N Y., vii, pi 31, fig 13 P 97 Tropidocaris alternata Beecher Ibid., fig 15 P 97 Tropidocaris alternata Beecher Ibid., fig Prestwichia randalli Beecher Eurypterus pennsylvanicus fig 18 C E 29, 1902, p 143, P 98 Hall 2nd Pa Geol Surv Rep pi 5, P 98 Thinopus antiquus Marsh From Morton, D 71, no 12, 1927, p 409 P 107 14 From Amer GeoL, Thinopus antiquus Marsh Ibid J Am Journ of Sc, n s PL 80, Vol 15, Hi-LL Amkr Pali-;ont Xo 58, Pl 59 INDEX TO VOLUME 15 Note: Figures refer to the continuous numbering of the volume (not of the separate bulletins); black face numerals refer to plates; light face, to pajfes A Actinopteria beta C alpha - Admete luffa Ambocoelia gregaria umbonata var gregaria Andicula n gen Anomalina affinis mantaensis wuellerstorfi Anomia ( ?) 46 46 206 207 Calathospongia 16 32 32 93 180 180 94 32 28 31 Camarotoechia allegania 4 Archinacella incerta Apedodus priscus Area saladoensis - 62 79 (?) samanensis Argyrotheca berryi wegemanni aequilateralis cancellatus celsus duplicatus 71 100 119 135 177 180 24 182 183 183 182 182 182 183 182 182 17 Asterigerina subacuta Athyris angelica lamellosa Aulostomella Aviculopecten aequilata 30 32 53 52,53 _ ellipticus patulus striatus tenuis 234 248 70 54 53 54 53 54 53 53 B Bellerophon maera Hoggsia n gen Bolivina applinae pisciformis Bonellita Kothriolepis minor Buliniina compressa inflata ovata 5 78 _ redfieldi contracta orbicularis 19 231 78 35 36 93 248 133 37 132 , sappho Cardium himertum Cassidulina subglobosa Caster, K E author of Bull 58 Cerithiopsis quemada Cerithium la(;visculum laevisculuni saladoense 35 33 33 33 restinense scitulus setigerus Cibicides alleni "I 40 14 12 12 americanus 21 dohertyi elongatus haidengeri lobatulus nucleata 20, 21 21 pseudoungerianus ungeriana 20 wuellerstorfi Cladodus carinatus see Bull 157 13 37 37 37 Chonetes lepidus 238 238 155 152 157 11 paytense Claiborne 68 73 74 carlli 145 87 81 82 81 84 149 149 150 29 136 30 137 30 137 31 137 31 31 248 formainifera, 50, p 57 Clathrospongia abacus Clavulina brayamensis bryamensis extans communis curta pallida parisiensis Cole W S., 73 238 21 128 34 34 34 34 pallida _ 5 author of 3i8 Bull 56 319 tumidum (loniophora cuivata ruedemanni (Jraniniysia 71 23f» 67 220 220 217 217 217 217 217 fi7a coniniunis fifi duplicata glabra 6() 86 66 66 subnasuta undata iiTOKularis (iiittulina 131 Gyi'acanthus sherwoodi Cyroceras stebos Cyroidina laevis 79 68 248 236 27 27 27 nitidula soldanii 4 H 86 16 8:" 21i) 246 78 77 78 24G 247 246 79 248 84 237 taylori Homacanthus acinaciforsubgen Hydnoceras nodosum 73 I n subgen 81 L aspera asperoides globosa marginata orbignyana caribaea semistriata strumosa _ _ 31 MO waverliensis Lingulina sp mesoneiisis 31 147 23 18 \'.'.] liingulodiscina pleurites 31 Lucina coibula _ Lyriopectcn alternatus 56 ^4 147 74 183 183 183 16 9;) rude - sulcata mytiliforme 44,45 45, 61 potens potens juvons ruedemanni depressum 31 fascial us 22 23 22 incsonensis raexicana Morgana costata ly 20 43 Mc>rrey, Margaret, co-author of Bull 55 85 Modiola 20 21 129 129 129 _ , _ crassata 11 18 gardnerae 7, _ 18 125 60 125 i:;i) 19 130 81 It 123 24 (.Mytilops) tclla convergens 12 If) of Misilus cornellina rotulata theta Meson formation, forams IS 11 _ M Mangclia jabonillnlensis Marginulina sp 5;) 19 19 19 58 42 42 44 43 43 n Levifusus iiuillicus Lingula cuyahoga Lenticulina oecidentalis novangliae gigas 31 scutella maxima 2!) _ Lagenonodosaria monilis Lagunites n gen Lepidocyclina 31 biton maidurii ventricosa vulgaris delia ligea alpha solox Lagena sulcata apiculata 125 125 187 196 198 186 191 195 192 193 195 193 193 187 197 187 187 196 188 188 92 147 147 147 mentor mentor mentor mentor mantor mentor umbcnaiuni 125 18 77 us pustulosus Iddingsia eornelli disparile mentoi' 18 18 43 45 46 43 42 15 43 43 44, 45 42 stephani see Orthothetes Hetcrcstegina antillae Holonenia rugosa Holoptychus american- n undosa Lci)(odesina bec-ki carinifera paralleluiii Harrisianella n gen Harrisianella peruviana Heliodus gibberulus Heniipronites crenistrea, mis Hopkinsiana I)arvula praocedens me59,63 23,38 Modiomorpha quadrula 64 rigida 64 chiMiiuiigcinsis Mytilarca 70 gibbosa 60, 62 lata 59 oecidentalis 60 59 regularis 215 214 21.^^ 215 213 214 214 214 213 320 32 lachrymosa stigmata 40, 41 onusta 40,41 167 169 speciosa 40 168 striatula 41a 168 Proihynchus angulatus _ 58 202 harrisi _ 44 202 nasutum 58 204 paeneplanus 44 203 quadratus 56, 61 201 Prothyris exuta 60 228 Protolimulus eriensis 76 241 - Pteriopecten suborbicu- laris 55 67 67 falcata lobata obsoleta perlata proto Salamanca sinuosa spatulata spio thalia Pulleniatina obliquiloculata Pulvinulina bryamensis canariensis umbonana _ _ angusta R Raphanulina Reticularia praematura Retzia sp Rhadinchthys sp Rhinoclavis Rhipidomella leucosia pennsylvanica Rhynchospirina scansa Rhynehospirina sp 24 181 156 32 32 78 248 37 38 84 150 150 32,34 181 136 34 25 26 Q Quinqueloculina nitidula soldanii 182 19 20 48, 49 quinqueloba taettowata Potalia advena canariensis ccuadorensis haidingeri 43 44 50 vanuxemi PuUenia bulloides _ 6 49 midwaycnsis 50 50 50 50 50 58 50 49 51 21 liLiraciii.i ungeriana 47,48,50 thetis trigonalis floforniis umbonata 46,49,51 _ -ii'ri.i 200 200 210 211 211 211 211 210 210 209 209 212 210 208 209 212 208 211 212 210 212 210 209 eudora galene gibbosa Pn' Robulus calcar deformis 183 Pteronites profundus rostratus Ptychopteria alata 47,48,49 beecheri 47,48,50 elongata eucrate 47,48,49 - _ 20 21 21 128 21 25 25 26 30 27 27 26 31 S Sagenodus 79 248 64 64 64 231 231 227 64 64 64 228 227 228 Schizophoria striatula 38 Schuchertella chemungensis 36 Sigmoilina celata Siphonina advena 21 Siphoninella bryamensis 20 Sphaeroidina bulloides chilostoma Sphenotus arcuatus 65 clavulus 63,65 contractus 63, 65 151 sp Sanguinolites eatus subtrun- undulatus Schizodus rhombeus chemungensis quadrancuneus gularis oblatus _ -.- palmerae 63 signatus 65 Spirifer allegheniensis 25 disjunctus 23,24 Spirodomus insignis 148 134 134 32 218 218 218 219 219 172 169 80 224 13 41 90 149 149 Spiroloculina calata Strombus buhrinus Strophalosia histricula murJcata 37 Stropheodonta demissa 36 Stylonurus beecheri 76 Surcula occidentalis 15 Syringothyris angulata 27,28,29 rajidalli 151 240 94 174 174 T Telescopium peruvianum Textularia candeiana flabelliformis 19 85 127 35 322 gramen 20 subhaueri Thinopus antiquus Thysanodictya expansa randalli Trilocullna angusta Tropidocaris alternata bicarinata interrupta Truncatulina alleni americana elongata - - 80 72 72 Turritella anceps negritoensis 249 238 238 Tylodictya warrenensis - _ 80 80 80 haidingeri lobatula nucleata pseudoungeriana tenera ungeriana wuellerstorfi — Turricula bonilla 127 128 16 239 239 239 29 136 137 80 137 81 137 26 31 31 100 12 74 78 79 238 U Uvigerina alata 6 38 38 138 39 133 39 39 11 11 5 19 73 74 33 37 133 beccari bryamensis 19 hispida niesonensis proboscidea pygmaea _, V Venericardia planicosta labraensis talara Vernuelina cyclostomata Virgulina bramletti compressa Woodsalia End of Volume XV n W subgen 79 ... Formation of Mexico By 58 11: S and Ruth Cole Gillespie Higher Fossil Faunas of the Upper Aileglieny Bv A'einiefl! E ( 'asler ^^,,.»-''-.|Hs,r„^,^^ BUI^IvETINS OF AMERICAN PAI.EONTOLOGY Vol 15... or definitions of genera are not agreed upon by recent authorities, the synonymies and descriptions of the genera are included The classification is that The age of the fauna of the senior author... described faunas : Upper Eocene of Texas, described by Cushman and Applin Lower Eocene of Texas, described by Mrs Plummer Upper Eocene of Mexico, described by Cole and Pliocene of Kar Nikobar, described