Bulletins of American paleontology (Bull. Am. paleontol.) Vol 542441968

EiECQ MUSEUM MARBIAt'klSTPRY SEP 11968 3% BULLETINS -**;*&&* OF AMERICAN PALEONTOLOGY (Founded 1895) Vol 54 No 244 GASTROPODS OF THE MIDDLE DEVONIAN ANDERDON LIMESTONE By Robert M Linsley 1968 Paleontological Research Institution Ithaca, New York 14850, U.S.A PALEONTOLOGIGAL RESEARCH INSTITUTION 1967 1968 - Kenneth President Vice-President Secretary-Treasurer E Caster William B Heroy Rebecca S Harris Katherine V W Palmer Director Armand Counsel Representative AAAS L Adams David Nicol Council Trustees Kenneth E Caster Donald W Fisher Katherine V W Palmer (Life) William B Heroy (1962-1968) (1966-1972) (1967-1973) Axel A Olsson (Life) Hans G Kugler (1963-1969) Rebecca S Harris (Life) Daniel B Sass (1965-1971) W Storrs Cole (1964-1970) BULLETINS OF AMERICAN PALEONTOLOGY and PALAEONTOGRAPHICAL AMERICANA Katherine V W Palmer, Editor Mrs Fay Briggs, Secretary Advisory Board Hans Kugler Jay Glenn Marks Kenneth E Caster A Myra Keen Axel A Olsson Complete titles and price of separate available list numbers may be had on application For reprint, Vols Kraus Reprint For reprint, vol I, Subscription may St., New Palaeontographica Corporation, 111 Fifth Ave., price of $16.00 per American Paleontology 1-23, Bulletins of Corp., 16 East 46th New see York, N.Y 10017, U.S.A Americana see Johnson Reprint York, N.Y 10003, U.S.A be entered at any time by volume or year, with average volume for Bulletins Numbers of Palaeontographica Amer- icana invoiced per issue Purchases in U.S.A for professional purposes deductible from income tax For sale by Paleontological Research Institution 109 Dearborn Place Ithaca, New York U.S.A 14850 are BULLETINS OF AMERICAN PALEONTOLOGY (Founded 1895) 54 Vol No 244 GASTROPODS OF THE MIDDLE DEVONIAN ANDERDON LIMESTONE By Robert M Linsley September 5, 1968 Paleontological Research Institution Ithaca, New York 14850, U.S.A Library oj Congress Catalog Card Number: Printed in the GS United State* of America 68-13i CON! VIS I PA(,F 333 Abstract General Statement • Stratigraphy of the Anderdon Limestone Detroit River (/roup — Lucas Dolomite 335 342 344 Dundee Limestone Detroit River 333 Group — Anderdon Stratigraphic correlation of the Limestone Anderdon Limestone 344 346 Occurrence of gastropods 351 Paleoecology 360 Preservation and preparation of material- 362 v s>-tematic 365 descriptions Bellerophontidae 365 Euomphalidae Kotomariidae 372 37X Lophospiridae 383 384 Raphistomatidae Portlockiellidae 394 Gosseletinidae 396 Platyceratidae 405 Microdomatidae +06 Anomphalidae 41 J'-Midophoridae 413 Xtritopsidae 414 Murchisoniidae 420 Palaeozygopleuridae 43 Streptacididae 438 Literature cited 438 Plate- 445 Text-figiris Aerial Drawing from geologic map aerial 340 view 345 Tables Numeral count Relative frequency of occurrence [species] per quarry of identifiable specimens 353 355 GASTROPODS OF THE MIDDLE DEVONIAN ANDERDON LIMESTONE Robert M Linsley ABSTRACT This paper is concerned with the stratigraphic occurrence and study of the large and interesting gastropod assemblage found in the Middle Devonian Anderdon Limestone This limestone is the uppermost unit of the largely calcareous Detroit River Group and its area of outcrop is apparently limited to southeastern Michigan, southwestern Ontario, and northwestern Ohio The overall stratigraphic position of the Anderdon Limestone is briefly discussed, though because of the limited value of gastropods as index fossils, this fauna provides little additional information regarding its relative age The comparative ages of the three gastropod beds of the Anderdon are discussed, and here suggested that they are not synchronous paleoecologicnl setting is also discussed briefly In general it was found that there are two primary ecologic environments represented within the -Anderdon One of these environments consists primarily of a biostromal aggregation of corals and the gastropods apparently lived in this protected environment The gastropods inhabiting this biostrome are predominantly small and this environment is represented in all three quarries that were visited by the author The second environment is known only from the Brunner, Mond Canada, Limited quarry of southeastern Ontario and consists of a fine calcilutite inhabited by an assemblage of gastropods whose average size greatly exceeds the average size of those found in the reef environment This second environment has been interpreted as being a protected back-reef or inter-reef environment Following the section on paleoecology there is a brief discussion of preservation and techniques Almost all of the gastropods occur as external molds in various states of preservation To study the gastropods it was necessary to make casts of each mold, and this was accomplished by using a latex molding compound Most photographs used in the plates are of the latex casts The bulk of this paper is devoted to systematic descriptions of new genera and species and to a brief discussion of other new- forms which are not well enough preserved to warrant description Over 50 different gastropods have been discovered in the Anderdon limestone, only two of which have previously been described The gastropods constitute a varied aggregation belonging to 27 genera representing the superfamilies Bellerophontacea, Macluritacea, Euomphalacea, Pleurotomariacea, Platyceratacea, Microdomatacea, Anomphalacea, Pseudophoracea, Neritacea, Murchisoniacea, Loxcnematacea, and Pyramidellacea Of the 50 species discussed, 33 are considered new Included in these 33 species are six genera which are comidered new They include Ehlprsina, Zalozone, Tylozonc, and Nodotirma (of the superfamily Pleurotomariacea), it is The (of the superfamily superfamily Murchisoniacea) Copidocatomus the Microdomatacea), and Crenulazona (of GENERAL STATEMENT This paper deals primarily with a discussion of the gastropod fauna of the Anderdon Limestone, the youngest formation of the Middle Devonian Detroit River Group exposed in southeastern Michigan, northwestern Ohio, and southeastern Ontario Gastropods comprise a remarkably large and distinctve element of the fauna of the Anderdon Limestone They seem limestone and hence are extremely useful to in be restricted to this differentiating the An- Bulletin 244 334 derdon Limestone from underlying formations of the Detroit River Group and the overlying Middle Devonian Dundee Limestone Over 50 different species of gastropods have been found in Limestone, but unfortunately many of these are unAnderdon the describable at the present time because of a lack of well-preserved specimens Nevertheless enough were specimens well-preserved and six new genera (Ehlersina, found to describe 33 new Crenulozona and Copidocatomus) Zalozone, Tylozone, Nodpnema, belonging to 27 different In all, 50 different species of gastropods species genera are discussed In addition to the formal descriptions of the species of gastro- pods of the Anderdon Limestone, a brief account of the stratigraphy of the Anderdon Formation is included, as are observations on the inferred ecology of the Anderdon sea Museum This study was initiated at the Paleontology of of when the author was a research last year of work by the author on the University of Michigan as- this During the study the research was sponsored by the National Science Foundation, and many of the expenses incurred in the final preparation of this manuscript were defrayed by giants from the Littauer Foundation and the Research Council of Colgate University Dr G M Ehlers of the University of Michigan suggested this sistant there project as a subject for a doctoral dissertation Throughout the entire period of study he proved to be a prime source of inspiration, and the author spent many stimulating hours discussing the various aspects of this problem with him Dr Ehlers also available not only the facilities of the University of Michigan seum of Paleontology but also provided the basic Anderdon gastropods that he had collected with the Dr E C Stumm and Dr R V Kesling, both of associated with the University of Michigan indebted to Dr Ehlers for his field trips to the The Dr of the Mu- collection of assistance of whom The author is are also further company and guidance on numerous outcrop areas of the Anderdon Limestone writer received and nature made many ideas concerning the possible extent Anderdon sea from Stumm Dr Kesling interesting discussions with also aided the progress of this study with numerous helpful observations on methodology and techniques Because of the small size of many of the gastropods, Dr Kesling's Anderdon Gastropods: Linsley familiarity 335 with micropaleontological techniques was particularly preparing and photographing the helpful in aiding the writer in specimens Mr Herbert Wienert, photographer for the University of Michigan Museum of Paleontology, helped the author with some of the photography The author greatly enjoyed the friendly and stimulating cor- respondence with Dr Ellis L Yochelson, Dr Roger L Batten, and Brooks Knight All three gentlemen gave unhesitheir time and greatly aided the determination of the tatingly of systematic position of many elements of this fauna, as well as the late Dr J fascinating suggestions regarding in many other problems encountered the course of this study Dr Henry Van der Schalie of the Museum University of Michigan aided the writer with of Zoology of the many informative hours of conversation regarding modern mollusks and thus pro- vided some possible analogies to the problems encountered in the Paleozoic snails Finally and most gratefully the author acknowledges the ungrudging Anne this hours E Linsley, manuscript ment and his JoAnn Hoehler wife, Linsley, and Mrs Haskell R Fenner spent They were their efforts are a in his many sister, preparing never failing source of encourage- sincerely appreciated STRATIGRAPHY OF THE ANDERON LIMESTONE To understand fully gastropods described in the this stratigraphic paper, it is distribution of the necessary to briefly de- Anderdon Limestone The term Anderdon Limestone was first used by W H Sherzer and A W Grabau (1908, p 408) for the "middle member" of the "Upper Monroe" strata, which, according to those authors, scribe the stratigraphy of the occupy a position between the underlying Sylvania Sandstone and the overlying Dundee Limestone No type locality or detailed deAnderdon Limestone was given by those investiga- scription of the tors (op cit.) In 1909 Sherzer and Grabau (1909, p 542) indicated that the term Anderdon Limestone was taken from the "Anderdon quarry, in Anderdon township, Essex County, Ontario, about miles east Bulletin 244 336 Amherstburg and of 15 miles south of Detroit." They (op cit.) de- Anderdon Limestone and the disconformity between this limestone and the overlying Dundee Limestone of the quarry, which is now owned by the Brunner, Mond Canada scribed the lithology of the Ltd In the text-figure and Grabau (1909, pp 541, 542, Anderdon Limestone overlies the reference, Sherzer indicated that the ) Rock Dolomite, which Flat is same respectivelv overlain They rests on the Sylvania Sandstone, and by the Amherstburg and Lucas Dolomites also described (op cit., Anderdon Limestone (not of the Detroit salt shaft, presented list of fossils cit., type Anderdon Limestone) (op cit., p 547) a combined from the "Anderdon bed" of the shaft and from the type Anderdon Limestone (op the occurrence of the pp 541, 542) a correlate of in Anderdon Township, and discussed pp 543, 544, 551-555) the correlation and faunal differ- ences of the Anderdon and other formations of the W H Upper Monroe W Grabau 1909, pp 553-556) presented a classification of the Monroe Formation of Michigan, Ohio, and western New York In this A C Lane, Charles S Prosser, Sherzer, and A ( classification the lower part of this "Lower Monroe graphic units, "formation" was designated the Bass Island series," composed of four strati- or Monroe" having a upper part the "Upper middle part the "Middle the single unit, the Sylvania Sandstone, Monroe," containing four units and the In ascending order the four divi- Detroit River Series were designated the Flat Rock Dolomite, Anderdon Limestone, Amherstburg Dolomite, and Lucas Dolomite sions of the Lane, Prosser, Sherzer, and Grabau (1909, p 555) stated that name Anderdon Limestone was " suggested by the Reverend " and "was adopted by Sherzer and Grabau Thomas Nattress the for the limestone exposed in the Anderdon quarry, Essex county, Ontario, two miles from Amherstburg, Ontario, and in reef the salt shaft at They m coral (1909, p Oakwood [part of Greater Detroit], Michigan." 553) also stated that the highest Silurian strata America are represented by the Monroe Formation, the Anderdon Limestone is a part of which The Anderdon Limestone was described in considerable detail and assigned to the Silurian System by W H Sherzer and A W Bull Amer Paleont., Vol 54 Plate 38 3C Anderdon Gastropods: Linsley Explanation of Plate 459 All figures of latex casts except figure 3c Figs 3a,h,c,4a,b,c,5a,c, X 2; figs la,2a,b,c,d,5b,6a,b,c, X 4; fig lb, X Fage Figure Pagodea falcatinoda Linsley, a,b n 406 sp UMMP 40384, showOblique basal view and side view of holotype, ing thin apertural lip, absence of an umbilicus, and sickle-shaped nodes at the periphery The lighting on fig lb emphasizes the growth lines, but fails to show the prominence of the middle, revolving lira Note carinate ornamentation of the neanic whorl 408 n gen., n sp views and oblique basal view of holotype, 40381 The primary light of fig 2a was placed at the lower right to emphasize whorl profile and sharp revolving carinae bordering the outer whorl face The primary light of fig 2c was placed at the lower left to emphasize growth lines, b Side view of paratype, 40383, showing sharp angulations bordering flattened outer whorl Copidocatomus coilarus Linsley, a,c,d Opposing UMMP side UMMP face 416 Naticopsis (Naticopsis) opimavoluta Linsley, n sp 40354 c Crossa,b Side view and apertural view of holotype, 40365 Note how the two shell layers section of paratype, thicken at the apertural shoulder and are separated from each other UMMP UMMP ( by solution ?) 417 Naticopsis (Naticopsis) planifrorts Linsley, n sp 40359, showa,c Oblique apical view and side view of holotype, ing sharp, deeply incised sutures, and the absence of a pronounced shoulder b,d Oblique apical view and side view of paratype, UMMP UMMP 40355 414 Naticopsis (Naticopsis) pegmihumerosa Linsley, n sp 40350, showing wide shoulders, a Apertural view of holotype, deep body whorl, and thickened aperture, b Oblique apical view of 40352 40360 c Apertural view of paratype, paratype, UMMP UMMP UMMP Turbonitella trunculinoda Linsley, a,b Side views of holotvpe, n UMMP UMMP 40368 sp 40367 c Side 418 view of paratype, Bulletin 244 460 Explanation of Plate 39 All figures of latex casts Figs la,b, X 2; figs 4a,e,g, X 8; all other figures X + Page Figure Omphalocirrus a,b 372 sp UMMP 40385, showBasal view and slightly oblique basal view of ing the spines around the periphery and the bundling of the growth lines Streptacis ? Side view of 438 sp UMMP 40544 Note fine costae and inclination of sutures 436 Palaeozygopleura joanni, Linsley, n sp Side view of paratype, UMMP 40480, showing extreme backward a obliquity of the costae of the ultimate whorl of this rotund individual, 40475 The ultimate whorl has been b Side view of paratype, broken back for about one-half of a volution, c Side view of para40476, showing smoothness of the nucleus and the erecttype, 40473 e ness of the earliest costae d Side view of paratype, 40481 Note the much finer costae on Side view of paratype, 40479 g Aperthe ultimate whorl, f Side view of paratype, 40478 h Side view of paratype, tural view of holotype, 40477 j,k Apertural 40482 i Side view of paratype, 40474, showing an extreme view and side view of paratype, condition of crowding of costae UMMP UMMP UMMP UMMP UMMP UMMP UMMP UMMP UMMP 436 Palaeozygopleura sibleyense Linsley, n sp Apertural view of paratype, UMMP 40471, showing wide pleural a angle of this immature specimen, b Side view of paratype, 40347, showing wide spacing of costae and slight constriction just 40470 d Side view below suture, c Side view of paratype, UMMP of holotype, UMMP UMMP 30686 Bull Amer Paleont., Vol 54 /i &$£A ' •* Plate 39 INDEX Note: The left hand bold face figures refer to the plates The right hand light figures refer to the pages A Index 32 Solenospira delphinuloides F 398 398 Euryzone Helicites Fagerstrom, Mich 333-350 341,351,361 Diodontopteria 373 dionysii, Straparollus 339 Disphyllum Detroit River Gr fragosus, Scalae- France Stone Company Quarry 35 339,341,344 Fritz, 351, 354, 356, 358, 359, 373- 376 Galloway, 423 334,337-338 340-350, 380 M 425 m anulata, - Erie Co., Ohio Euconospira Euomphalus sp sp sp A B C Euryzone H hamiltoniae, Loxonema 383 387 339 347, 348 380 354,356,376378, 403 36 376 36 376 36 377 338,351,353, A 30,31 B 31 Helicites Helix Heterophrentis Homotoma Hormotoma Hormotomina imbricata, Murchisonia 436,437 398 378 339 424 433 349,428-431 411 incomptum, Ceri- 404 402 403 Eversole Chart Zone 436, 437 Palaeozygopleura 355, 362, 397?sp ?sp 28 341,344,353 355, 358, 359 394-396, 413 30 Emmonsia No- donema 353, 355, 379, 380, 383 sp 33 353, 355, 424 341,351,361 333,334,338, topteria Ehlersina ? Mur- chisonia Ectomaria eliana, Luciella 372 384 335-337,40941 1-, 422-423 W qracilicrista, E Diodon- Ompha- Goniospira Grabau, A 344, 346-350 403, 404 ehlersi, 350 351 J J goldfussi locirrus 334-338,341, G 350 M A Germany 434 367 Ls 342-346,351361 363 G 33 354,356,384 Donaldina 37 354.356,413414 trochus diversiformis, Straparollus (Serpulo- Donaldiella 38 354,356,406- 408 Coelozone 29 353, 356, 397 339.341,351 Favosites 335 Fenner, Mrs H R 384 filosa, Goniospira 336,338 Flat Rock Dol 426 336,348 Murchisonia Ehlers, 349 fasciata, desiderata, Donax Dundee A Pagodea 33 353, 355, 427 428 411 depressum, Isonema Murchisonia spira) J falcatinoda, doludisubzona, Detroit 421-423 33 354,356,434, 435 347 thioides Ingersoll, Ont Isochilina 339 349,411-413 Isonema 411 extenuata, Ectomaria Murchisonia J 423 422,423 James Bay, Canada 462 349.350 Index j Murchisonia anus, Mesocoelia 433 Murchisonia 433 Jedria 416 joanni, Palaeozygo39 344,354,356, pleura 436, 437 K 334.338.340 342-348 338,339 335,364,384, Kloedenia Knight, J B 394, 395, 410, 411 Kwataboahegan 350 River L lacunata, Zalozone 26, 27, 28 353, 355, 384- 388 336 Lane, A C latitornata, Euryzone 29, 30 353, 355, 358, 399-401 Leiorhynchus LeRoy, N Y 338 347 lindstromi, Scalaetrochus 413,414 413 335 335 Trochus Linsley, A E Linsley, JoAnn linsleyi, Murchi- (Hormotomina) Lodonaria sonia Loxonema Loxoplocus 430 431 436,437 33 354, 356, 383, 384 Lucas Dol 336-338,340, 342, 343, 345- 351 Luciella 385,387 M Murchisonia (Hormotomina) niaia, manitobensis, 428-430 Ompha372 349,350 428,433 434 333,334,337 locirrus Martinson, N W Mesocoelia Michelia Michigan Microdoma minuta, Murchisonia Monroe Fm Mourlonia ?sp 434 Murchisonia (Hormotomina) 31 409-411 421 336, 337 353,355,378 378 430 N Naticopsis V Kesling, R 344, 351, 409, 419-430, 433, Index 354, 356, 434, Sterptacis phillipsii, 438 438 414-417 Naticopsis 39 ?sp planifrons, 38 354, 356, 415- Naticopsis 418 Stringocephalus Stumm, Zone 351 334, 338, 340- E C 348,371 354, 356, 405, Platyceras 37 sp Pleurotomaria verna Pleurotrochus Portlockiella Prosser, C S Punctoprimitia 406 405 397 409 394 336 339 stummi, Bellerophon 25 358,353,355 371 418 subcostata, Naticopsis 415 subovatus, Naticopsis 335-338,345, Svlvania Ss 348 339,350 Syringostroma 385, 389 394, 395 Raphistoma Rhineoderma S Sandusky Co., Scalaetrochus Scalites serpula, Ohio Euomphalus Serpularia Serpulospira Shansiella Sherzer, W H Sibley Quarry 347,348 413 389 374 341,342,351- sibleyense, Palaeozygopleura telescopium, Cerithioides 339,341,351, 361 39 341,354,356, 358, 436, 437 trepomena, Murchisonia 32 341, 353, 355, 358, 359, 425- Silica Silica, Fm Ohio , 427 346 342.343,346 348 Solenospira extenuatum Stauffer, C R 421-423 337-339,347 Stictostroma anderdonense Straparella Straparollus Straparollus (Euomphalus) Straparollus (Serpulospira) 339 412 373-378,126 420 tricarinata Homotoma Hormotoma Microdoma 424 424 36 344,358,354, 356, 423 tricarinatus, 409,423 Pleurotrochus tricincta, 420 Murchisonia Trochus 413 358,365-371 (Scalaetrochus) Tropidodiscus tropidophora, Loxoplocus Murchisonia 354,356,383, 384 383 Irunculinoda, Turbonitella 38 354,356,418, sibleyensis, Murchisonia 434,435 416 380 Trachydoma Trechmannia 394 335, 336 394 Tapinotomaria 373 373-376 361, 363, 368, 370, 372, 411 Conocardium T 348 Raisin River Dol 419 turbinatus, 419 416,418,419 Murchisonia Turbonitella turricula, Tylozone 26 353, 355, 392, 393 Tylozone typa, 334, 338, 353, 355, 388-393 Caiaschisma Van der Schalie, H 354, 356, 376378, 403 vasulites, 339, 341, 344, ventricosa, verna, 464 335 Bellerophon 371 Naticopsis 415 Coelozone 351,373 404 397 Index Pleurotomaria 397 vesculilineatus, 25 344,353.355, 365-370 Tropidodiscus vesna, Palaeozygo- vittatus, Donax Wienert, H Williams, M Y Wilson, A E 438 335 337 349 Yochelson, E L 335, 364 pleura vetusta, Platyceras WhitfielcU, Streptacis 436 405 367 W Waines, R Wenz, W H 350 384 Zalozone 465 334, 338, 353, 355, 384-388 XL (No 184) 996 pp., pis 16.00 Type and 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Institution 109 Dearborn Place Ithaca, New York U.S.A 14850 are BULLETINS OF AMERICAN PALEONTOLOGY (Founded 1895) 54 Vol No 244 GASTROPODS OF THE MIDDLE DEVONIAN ANDERDON LIMESTONE By Robert M Linsley... species of gastropods species genera are discussed In addition to the formal descriptions of the species of gastro- pods of the Anderdon Limestone, a brief account of the stratigraphy of the Anderdon