ON THE ORIGIN OF SPECIES, DARWIN

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ON THE ORIGIN OF SPECIES, DARWIN

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ON THE ORIGIN OF SPECIES BY MEANS OF NATURAL SELECTION, PRESERVATION OF FAVOURED RACES IN THE STRUGGLE FOR LIFE By CHARLES DARWIN, FELLOW OF THE ROYAL, GEOLOGICAL, LINN^iAN, AUTHOR OF JOURNAL OF RESEARCHES DURING ROUND THE WORLD.' ' M.A., ETC., SOCIETIES; H M S BEAGLE'S VOYAGE LONDON: JOHN MURRAY, ALBEMARLE STREET 1859 The r : rjlit, nf Translation, is reserved LONDON: PRINTED BT W CLOWES AND SONS, STAMFORD AND CHARING CROSS STKISKT, 3fc5 CONTENTS Introduction Page CHAPTER I Variation under Domestication — Correlation of Growth — — of distinguishing between Varieties and Species — Origin of Domestic Varieties from one or more Species — Domestic Pigeons, Differences and Origin — Principle of Selection anciently followed, — Methodical and Unconscious Selection — Unknown Origin of our Domestic Productions — Circumstances favourable — — Character Causes of Variability Inheritance Effects of Habit of Domestic Varieties Difficulty their its Effects to Man's power of Selection 7-43 CHAPTER II Variation under Nature Variability — Individual ranging, much — — — Wide Doubtful species and common species vary most Spegenera in any country vary more than the species differences diffused, cies of the larger — of the smaller genera Many of the species of the larger genera resemble varieties in being very closely, but unequally, related 44-59 to each other, and in having restricted ranges CONTENTS VI CHAPTEE III Struggle for Existence — — The term used in a wide sense Geopowers of increase Bapid increase of naturalised animals and plants Nature of the checks to increase Competition universal Effects of climate Protection from the number of individuals Complex relations of all animals and plants throughout nature Struggle for life most severe between individuals and varieties of the same species often severe between species of the same genus The relation of organism to organism the most important of all relations Page 60-79 Bears on natural selection metrical — — — — — — — ; — CHAPTEE IV Natural Selection Natural Selection — — its power compared with man's selection its its power at all ages power on characters of trifling importance Sexual Selection On the generality of interand on both sexes -Circumstances crosses between individuals of the same species favourable and unfavourable to Natural Selection, namely, Slow action intercrossing, isolation, number of individuals Extinction caused by Natural Selection Divergence of Character, related to the diversity of inhabitants of any small area, Action of Natural Selection, through and to naturalisation Divergence of Character and Extinction, on the descendants Explains the Grouping of all organic from a common parent 80-130 beings • — — — — — — — — — CHAPTEE V Laws of Variation — Use and combined with — Acclimatisation — Correlation of growth — Compensation and economy of growth — False correlations — Multiple, rudimentary, and lowly organised structures variable — Parts developed in an unusual manner are highly variable characters more variable than generic secondary sexual characters variable — Species of the same genus vary in an analogous manner — Keversions to 131-170 long-lost characters — Summary conditions Effects of external natural selection ; disuse, organs of flight and of vision : : specific — CONTENTS Vll CHAPTER VI Difficulties on Theory — — on the theory of descent with modification TransitionsAbsence or rarity of transitional varieties Transitions in habits Species with Diversified habits in the same species of life Organs of habits widely different from those of their allies extreme perfection Means of transition Cases of difficulty Xatura non facit saltum Organs of small importance Organs not in all cases absolutely perfect The law of Unity of Type and of the Conditions of Existence embraced by the theory of Page 171-206 Natural Selection Difficulties — — — — — — — — CHAPTER VII Instinct — — Aphides and ants — Instincts variable — — Natural of the cuckoo, bees — Slave-making ants — Hive-bee, — on the theory of the Natural — Neuter or — Summary Instincts comparable with habits, but different in their origin Instincts graduated Domestic instincts, their origin ostrich, and parasitic cell-making instinct instincts its Difficulties Selection of instincts sterile insects 207-244 CHAPTER VIII Hybridism Distinction between the sterility of first crosses — and of hybrids by close inter- Sterility various in degree, not universal, affected breeding, removed — by domestication — Laws governing the sterility endowment, but incidental on other differences Causes of the sterility of first crosses and of hybrids Parallelism between the effects of changed conditions of life and crossing Fertility of varieties when crossed and of their mongrel offspring not universal Hybrids and mongrels compared independently of their fertility Summary 245-278 of hybrids — Sterility not a special — — — — CONTENTS Vlll CHAPTEE IX On the Imperfection of the Geological Record On the absence of intermediate varieties at the present day the nature of extinct intermediate varieties On ; — On — on their number the vast lapse of time, as inferred from the rate of deposi- — On the poorness of our paheontological — in any one formation On the absence of intermediate — On — On the sudden appearance of groups of tion and of denudation collections — On the intermittence of geological formations varieties species sudden appearance in the lowest known their fossiliferous strata Page 279-311 CHAPTEE On the Geological On X Succession of Organic Beings / the slow and successive appearance of change different rates of — Species once — On their — rules in their appear— On Extinction — new Groups of species follow the same general lost species not reappear ance and disappearance as single species On simultaneous changes in the forms of world — On the — On the state of development On the succession of the same types within Summary of preceding and present chapters living species life affinities of extinct species to throughout the each other and to of ancient forms the same areas — — 312-345 CHAPTEE XL Geographical Distribution Present distribution cannot be accounted for by differences in phyImportance of barriers' Affinity of the prosical conditions — ductions of the same continent — — Centres of creation — Means by changes of climate and of the level of the land, and by occasional means Dispersal during the Glacial period 346-382 co-extensive with the world of dispersal, — CONTENTS CHAPTEE IX XII —continued productions — On the inhabitants Geographical Distribution Distribution of fresh-water of — Absence of Batrachians and of Mamthose of mals — On the of the inhabitants of islands the nearest mainland — On colonisation from the nearest source and prewith subsequent modification — Summary of the oceanic islands terrestrial to relation last Page 383-410 sent chapters CHAPTEE Mutual XIII Affinities of Organic Beings Morphology Embryology Rudimentary Organs : : : — Natural system — descent modification — of — Descent — Analogical or adaptive characters always used — complex and radiating — Extinction parates and groups — Morphology, between members of the same between parts of the same individual — Classification, groups subordinate to groups Bules and difficulties in classification, 'with explained on the theory of Classification varieties in classification Affinities, general, se- defines class, Embryology, laws of, explained by variations not supervening at an early age, and being inherited at a corresponding age Rudimentary organs their origin explained Summary 411-458 — — ; CHAPTEE XIV Recapitulation and Conclusion Recapitulation of the difficulties on the theory of Natural Selection Recapitulation of the general and special circumstances in its — — Caifses of the general in the immutability of — How the theory of natural selection may be adoption on the study of Natural extended — Effects of 459-490 history — Concluding remarks belief favour species far its Index 491 INSTRUCTION TO BINDER The Diagram to front page 117, and to Volume face the latter part of the ON THE ORIGIN OF SPECIES INTRODUCTION When much on board H.M.S ' Beagle/ as naturalist, I was struck with certain facts in the distribution of the inhabitants of South America, and in the geological relations of the present to the past inhabitants of that con- These facts seemed to me to throw some light on the origin of species that mystery of mysteries, as it has been called by one of our greatest philosophers On my return home, it occurred to me, in 1837, that something might perhaps be made out on this question by patiently accumulating and reflecting on all sorts of facts which could possibly have any bearing on it After five years' work I allowed myself to speculate on the subject, and drew up some short notes these I enlarged in 1844 into a sketch of the conclusions, which then seemed to me probable from that period to the present day I have steadily pursued the same object I hope that I may be excused for entering on these personal details, as I give them to show that I have not been hasty in coming to a decision tinent — ; : My work is now Dearly finished but as it will take two or three more years to complete it, and as my health is far from strong, I have been urged to publish this Abstract I have more especially been induced to this, as Mr Wallace, who is now studying the B me ; INTRODUCTION natural history of the at almost exactly the Malay archipelago, has arrived same general conclusions that I have on the origin of species Last year he sent to me a memoir on this subject, with a request that I would forward it who to Sir Charles Lyell, nean Society, and it is sent it to the Lin- published in the third volume of the Journal of that Society Sir C Lyell and Dr — Hooker, who both knew of my work the latter having read my sketch of 1844 honoured me by thinking it advisable to publish, with Mr Wallace's excellent memoir, some brief extracts from my manuscripts This Abstract, which I now publish, must necessarily be imperfect I cannot here give references and autho- — rities for my several statements ; and I must the reader reposing some confidence in No doubt errors will have crept in, my trust to accuracy though I hope I have always been cautious in trusting to good authorities alone I can here give only the general conclusions at which I have arrived, with a few facts in illustration, but which, I hope, in most cases will can feel more suffice No one sensible than I of the necessity of here- after publishing in detail all the facts, with references, on which my conclusions have been grounded and I hope in a future work to this For I am well aware that scarcely a single point is discussed in this volume on which facts cannot be adduced, often apparently leading to conclusions directly opposite to those at which A fair result can be obtained only by I have arrived fully stating and balancing the facts and arguments on both sides of each question and this cannot possibly be ; ; here done I much regret that want of space prevents my having the satisfaction of acknowledging the generous assistance which I have received from very many naturalists, some of them personally unknown to me I cannot, however, OS! 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