... for crop improvement Trends Biotechnol 26: 531–537 Chapter Molecular Biology of Secondary Metabolism: Case Study for Glycyrrhiza Plants Hiroaki Hayashi Abstract Licorice (roots and stolons of ... al., 2003) Molecular Biology of Secondary Metabolism 99 the high-level accumulation (more than 2% of dry weight) of soyasaponins (Hayashi et al., 2003) Furthermore, enzyme activity of UDP-glucuronic ... condensation of two molecules of farnesyl diphosphate into squalene, a common precursor of sterols and Molecular Biology of Secondary Metabolism mevalonic acid 97 farnesyl diphosphate SQS squalene O 2,3-oxidosqualene...