... lines The phenotypic variance of the trait in the i th line is V and ; , the mean and the variance of the distribution of the variances of the lines are V and V(1; ), respectively In each line, ... to the evolution of the between- and within-line variance of a quantitative trait In such analyses, the between-line variance, in the absence of maternal effects, is essentially genetic, although ... environmental variance between the lines, the variances of the phenotypic and additive variances of the lines will be the same and, therefore: where fl j CV(V and h are the coefficient of variation of the...