... of ADH4 in the mucus-secreting cells of pylorus(Fig. 3G). Therefore, in the surface epithelium, ADH1 andADH4 only overlapped in the gastric region close to the pylorus. The endothelium lining ... throughout the epithelium in intestinal villi andcrypts of Lieberku¨hn (Fig. 4A,B). By IHC, ADH4 wasprominent in the epithelial cells lining intestinal villi, in contrast to crypts of Lieberku¨hn, ... Both enzymes were detected in mucosabut, in general, ADH4 was found in outer cell layers, lining the lumen, while ADH1 was detected in the inner cell layers. Of interest were the sharp discontinuities...